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Vascular Plants of California
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Notholaena californica
CALIFORNIA CLOAK-FERN


Higher Taxonomy
Family: PteridaceaeView DescriptionDichotomous Key
Common Name: BRAKE FAMILY
Habit: Perennial herb, in soil or on or among rocks; rhizome creeping to erect, scaly. Leaf: generally all +- alike (or of 2 kinds, fertile, sterile), generally < 50 cm, often < 25 cm; stipe generally thin, wiry, often dark, ×-section with vascular strands generally 1--3, less often many in circle; blade generally pinnate or +- palmate-pinnate (see Adiantum), often >= 2-compound, abaxially often with glands, +- powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or other, veins generally free. Sporangia: in sori or not, marginal, submarginal, or along veins, covered by recurved, often modified segment margins (false indusia) or not; true indusia 0; spores spheric, sides flat or not, scar with 3 radiating branches.
Genera In Family: +- 40 genera, 500 species: worldwide, especially dry areas. Note: CA members of Cheilanthes moved to the distantly related Myriopteris; Pellaea breweri to be moved as well, from a to-be-redefined Pellaea; traditional, often untenable limits of genera outside CA also being clarified using molecular phylogenetics.
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith & Thomas Lemieux, except as noted
Scientific Editor: Alan R. Smith, Thomas J. Rosatti.
Genus: NotholaenaView Description 


Habit: Plant in soil or granite rock crevices; rhizome short-creeping to +- erect, scales lance-linear. Leaf: stipe generally cylindric, dark brown to black, glabrous to +- scaly; blade 2--4-pinnate, segments generally sessile, +- narrower at base or not. Sporangia: in +- continuous, marginal bands; segment margin recurved, partly covering sporangia, unmodified; spores finely ridged or granular, often +- black.
Species In Genus: 34 species: generally Mexico, southwestern United States, few in Caribbean, South America. Etymology: (Greek: false cloak, from leaf blade margin not reflexed as it is in Cheilanthes)
eFlora Treatment Author: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson
Unabridged Reference: Tryon 1956 Contr Gray Herb 179:1--106
Notholaena californica D.C. Eaton
NATIVE
Habit: Rhizome scales rigid, with +- black midrib +- to margins, finely ciliate. Leaf: 3-pinnate, +- 3--13 cm; blade axes brown to black, glabrous or with white to yellow exudate; lowermost pinnae each more developed on basal side; segment abaxially covered with white to yellow exudate, hairs 0, scales 0, adaxially sparsely dotted with white to yellow exudate. Sporangia: 32-spored. Chromosomes: n=2n=150.
Ecology: Dry rocky slopes, rock crevices, under rock ledges; Elevation: 200--1300 m. Bioregional Distribution: SCo, s ChI, TR, PR, DSon; Distribution Outside California: Arizona, northwestern Mexico. Note: Apogamous. At least 2 entities in California chemically distinct: 1 with pale to bright yellow exudate on leaf abaxially (Notholaena californica subsp. californica), 1 with white exudate on leaf abaxially (Notholaena californica subsp. leucophylla Windham). Gene flow between them where they overlap geographically (s California) 0 due to apogamy, and pentaploids in Arizona suggest further study needed to decide if taxonomic recognition warranted.
Synonyms: Notholaena candida (M. Martens & Galeotti) Hook. var. accessita Jeps.
Jepson eFlora Author: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson
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Botanical illustration including Notholaena californica

botanical illustration including Notholaena californica

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Citation for this treatment: Ruth E.B. Kirkpatrick, Alan R. Smith, Thomas Lemieux & Edward Alverson 2012, Notholaena californica, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=34721, accessed on November 18, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora, http://ucjeps.berkeley.edu/eflora/, accessed on November 18, 2019.

Notholaena californica subsp. californica
click for enlargement
© 2012 Keir Morse
Notholaena californica subsp. californica
click for enlargement
© 2012 Keir Morse
Notholaena californica subsp. californica
click for enlargement
© 2012 Keir Morse
Notholaena californica subsp. californica
click for enlargement
© 2012 Keir Morse
Notholaena californica subsp. californica
click for enlargement
© 2012 Keir Morse

More photos of Notholaena californica subsp. californica in CalPhotos



Geographic subdivisions for Notholaena californica:
SCo, s ChI, TR, PR, DSon
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.