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Nemacladus pinnatifidus

Higher Taxonomy
Family: CampanulaceaeView DescriptionDichotomous Key

Habit: Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, +- fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or +- fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1--3, placentas axile or parietal, ovules many, style generally 1, 2--5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
Genera In Family: +- 90 genera, +- 2500 species: worldwide. Note: Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus.
eFlora Treatment Author: Nancy R. Morin, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: NemacladusView DescriptionDichotomous Key

Habit: Annual, from taproot. Stem: prostrate, decumbent, or erect; base generally +- brown or +- purple; branches 0 or below middle. Leaf: basal; petiole short or 0. Inflorescence: +- raceme-like; bract 1 per flower, small; pedicel generally thread-like. Flower: inverted or not; sepals linear to triangular; corolla +- radial or 2-lipped, lobes 5; filaments free at base, fused into tube around style distally, appendages attached to a stalk or directly on 2 adjacent filaments, each with 2--12 cells, anthers free, all alike; ovary superior to 1/2 inferior, hemispheric to obconic, nectary glands 3, mounded or donut-like, on free part of ovary, stigma 2-lobed, papillate. Fruit: generally > hypanthium, hemispheric to fusiform, top pointed or rounded, chambers 2; open at top generally by 2 valves (or circumscissile). Seed: elliptic to oblong.
Species In Genus: 18 species: southwestern United States, northwestern Mexico. Etymology: (Greek: thread-like branch) Note: In descriptions, "filaments" including both free and fused parts thereof.
Unabridged Note: Taxonomic changes from TJM (1993) based on ITS, atpB, morphology. Parishella californica nested within Nemacladus, in which it is here treated.

Nemacladus pinnatifidus Greene
Habit: Erect, 6--20 cm, branches many from base to below middle. Stem: stiffly erect. Leaf: 5--20 mm, oblanceolate, deeply pinnately lobed, toothed or entire, glabrous, narrowed to a long petiole. Inflorescence: axis +- straight; bracts 2--5 mm, linear to elliptic, appressed to pedicel base, proximal pinnately lobed; pedicels 5--15 mm, < 0.1 mm diam, ascending to spreading, straight, tip hooked upward abruptly. Flower: not inverted; hypanthium 0.5--0.7 mm; sepals 1--1.2 mm, linear to narrowly triangular, separated by a sinus, erect; corolla not conspicuously 2-lipped, divided +- 1/3 to base, glabrous, white or faintly pink-tinged, tube bell-shaped, 1--1.4 mm, lobes elliptic, 0.5--0.8 mm, adaxial 2 reflexed, 3 abaxial spreading, or all reflexed; filaments arched, +- 0.7 mm, tip strongly curved, glabrous, appendage continuous with filament base, cells slender, acute, anthers 0.1--0.2 mm; ovary 1/2-inferior in fruit. Fruit: 3--4 mm, base, tip acute. Seed: 0.5 mm, elliptic, with widely spaced zigzag ridges.
Ecology: Dry washes, burned areas, chaparral; Elevation: 300--1520 m. Bioregional Distribution: SCo, SnGb, PR, DSon; Distribution Outside California: Baja California. Flowering Time: May--Jun
eFlora Treatment Author: Nancy R. Morin
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botanical illustration including Nemacladus pinnatifidus


Citation for this treatment: Nancy R. Morin 2016. Nemacladus pinnatifidus, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 05, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on May 05, 2016.

Nemacladus pinnatifidus
click for enlargement
© 2010 Steve Matson
Nemacladus pinnatifidus
click for enlargement
© 2010 Steve Matson
Nemacladus pinnatifidus
click for enlargement
© 2010 Steve Matson
Nemacladus pinnatifidus
click for enlargement
© 2010 Steve Matson
Nemacladus pinnatifidus
click for enlargement
© 2010 Steve Matson

More photos of Nemacladus pinnatifidus in CalPhotos

Geographic subdivisions for Nemacladus pinnatifidus:
SCo, SnGb, PR, DSon;
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.