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Nemacladus montanus

Higher Taxonomy
Family: CampanulaceaeView DescriptionDichotomous Key

Habit: Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, +- fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or +- fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1--3, placentas axile or parietal, ovules many, style generally 1, 2--5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
Genera In Family: +- 90 genera, +- 2500 species: worldwide. Note: Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus.
eFlora Treatment Author: Nancy R. Morin, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: NemacladusView DescriptionDichotomous Key

Habit: Annual, from taproot. Stem: prostrate, decumbent, or erect; base generally +- brown or +- purple; branches 0 or below middle. Leaf: basal; petiole short or 0. Inflorescence: +- raceme-like; bract 1 per flower, small; pedicel generally thread-like. Flower: inverted or not; sepals linear to triangular; corolla +- radial or 2-lipped, lobes 5; filaments free at base, fused into tube around style distally, appendages attached to a stalk or directly on 2 adjacent filaments, each with 2--12 cells, anthers free, all alike; ovary superior to 1/2 inferior, hemispheric to obconic, nectary glands 3, mounded or donut-like, on free part of ovary, stigma 2-lobed, papillate. Fruit: generally > hypanthium, hemispheric to fusiform, top pointed or rounded, chambers 2; open at top generally by 2 valves (or circumscissile). Seed: elliptic to oblong.
Species In Genus: 18 species: southwestern United States, northwestern Mexico. Etymology: (Greek: thread-like branch) Note: In descriptions, "filaments" including both free and fused parts thereof.
Unabridged Note: Taxonomic changes from TJM (1993) based on ITS, atpB, morphology. Parishella californica nested within Nemacladus, in which it is here treated.

Nemacladus montanus Greene
Habit: Erect, 8--18 cm, 1st branches generally 2--2.5 cm from base. Stem: ascending. Leaf: 6--18 mm, oblanceolate to elliptic, entire or obscurely toothed, hairy, narrowed to short, wide petiole. Inflorescence: axis strongly zigzag; bracts 1--3 mm, linear to lanceolate, spreading; pedicels 10--15 mm, 0.1 mm diam, ascending, generally straight, tip upturned. Flower: not inverted; hypanthium +- 1 mm; sepals +- 2 mm, triangular, erect; corolla conspicuously 2-lipped, divided +- to base, white, glabrous or ciliate, 3 adaxial lobes erect, 2 mm, yellow spot at base and 2 maroon arches above, 2 abaxial spreading, 3--4 mm, 2 maroon lines near base; filaments declined, +- 3 mm, tip +- curved, glabrous, appendage pad at base of filaments, cells obtuse, anthers +- 0.5 mm; ovary 1/2-inferior. Fruit: 2.5--3 mm, base acute, tip +- pointed. Seed: +- 1 mm, elliptic, with deeply impressed, vertical lines. Chromosomes: n=9.
Ecology: Serpentine soils; Elevation: 300--1200 m. Bioregional Distribution: NCoRI, NCoRH, n ScV, se SnFrB (Mount Hamilton). Flowering Time: May--Jul
eFlora Treatment Author: Nancy R. Morin
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Citation for this treatment: Nancy R. Morin 2016. Nemacladus montanus, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on February 11, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on February 11, 2016.

Nemacladus montanus
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© 2011 Aaron Schusteff
Nemacladus montanus
click for enlargement
© 2011 Aaron Schusteff
Nemacladus montanus
click for enlargement
© 2011 Aaron Schusteff
Nemacladus montanus
click for enlargement
© 2011 Aaron Schusteff
Nemacladus montanus
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© 2012 Aaron Schusteff
Nemacladus montanus
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© 2010 Steve Matson

More photos of Nemacladus montanus in CalPhotos

Geographic subdivisions for Nemacladus montanus:
NCoRI, NCoRH, n ScV, se SnFrB (Mount Hamilton).
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.