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    Vascular Plants of California
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Lycopus asper

Higher Taxonomy
Family: Lamiaceae (Labiatae)View DescriptionDichotomous Key

Common Name: MINT FAMILY
Habit: Annual to shrub [tree, vine], glabrous to hairy, generally aromatic. Stem: generally erect, generally 4-angled. Leaf: generally simple to deeply lobed, generally opposite, generally gland-dotted. Inflorescence: generally cymes, generally many in dense axillary clusters surrounding stem, generally separated by evident internodes or collectively crowded, spike- or panicle-like, occasionally head-like or raceme, subtended by leaves or bracts; flowers sessile or pedicelled. Flower: generally bisexual; calyx generally 5-lobed, radial to bilateral; corolla generally bilateral, 1--2-lipped, upper lip entire or 2-lobed, +- flat to hood-like, occasionally 0, lower lip generally 3-lobed; stamens generally 4, epipetalous, generally exserted, paired, pairs generally unequal, occasionally 2, staminodes 2 or 0; ovary superior, generally 4-lobed to base chambers 2, ovules 2 per chamber, style 1, generally arising from center at junction of lobes, stigmas generally 2. Fruit: generally 4 nutlets, generally ovoid to oblong, smooth.
Genera In Family: +- 230 genera, 7200 species: worldwide. Many cultivated for herbs, oils (Lavandula, lavender; Mentha, mint; Rosmarinus, rosemary; Thymus, thyme), some cultivated as ornamental (in California Cedronella, Leonotis, Monarda, Phlomis). Note: Moluccella laevis L., shell flower, historical waif in California. Satureja calamintha (L.) Scheele subsp. ascendens (Jordan) Briq. reported as alien but not naturalized. Salazaria moved to Scutellaria; California Hyptis moved to Condea, California Satureja moved to Clinopodium.
eFlora Treatment Author: Dieter H. Wilken & Margriet Wetherwax, family description, key to genera, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.
Genus: LycopusView DescriptionDichotomous Key

Common Name: BUGLEWEED
Habit: Perennial herb from rhizome, glabrous or hairy. Stem: erect, branched or not. Leaf: short-petioled to sessile; blade generally ovate to lanceolate, margin toothed to deeply lobed or cut. Inflorescence: head-like in leaf axils. Flower: calyx generally 5-lobed, lobes +- equal, obtuse to short-awned; corolla +- bilateral, not 2-lipped, generally 4-lobed, lobes +- unequal; stamens 2, exserted, staminodes 2, minute, club-shaped; style exserted. Fruit: nutlets +- compressed, truncate to rounded, edges corky-thickened.
Species In Genus: 14 species: temperate North America, Eurasia, Australia. Etymology: (Greek: wolf foot, from French common name)
eFlora Treatment Author: John M. Miller & Dieter H. Wilken
Reference: Moon et al. 2006 J Pl Res 119:633--644
Unabridged Reference: Moon & Hong 2003 Ann Bot Fenn 40:191--198; Henderson 1962 Amer Midl Naturalist 68:95--135

Lycopus asper Greene
Habit: Rhizome tips thicker, tuber-like. Stem: erect, 3--8(10) dm, puberulent to short-hairy. Leaf: +- sessile, 2.5--7(9) cm, lanceolate to narrowly elliptic, serrate, glabrous to puberulent. Flower: calyx lobes awl-like, acuminate to short-awned; corolla 3--5 mm, +- > calyx, white. Fruit: nutlet 1.5--2 mm; tip +- truncate, occasionally minutely toothed. Chromosomes: 2n=22.
Ecology: Moist areas, marshes, streambanks; Elevation: < 1400 m. Bioregional Distribution: CA-FP, GB; Distribution Outside California: to western Canada, Great Plains. Flowering Time: Jun--Oct
Unabridged Synonyms: Lycopus lucidus Turcz. ex Benth., misappl.
eFlora Treatment Author: John M. Miller & Dieter H. Wilken
Reference: Moon et al. 2006 J Pl Res 119:633--644
Unabridged Reference: Moon & Hong 2003 Ann Bot Fenn 40:191--198; Henderson 1962 Amer Midl Naturalist 68:95--135
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botanical illustration including Lycopus asper


Citation for this treatment: John M. Miller & Dieter H. Wilken 2012, Lycopus asper, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on July 19, 2018.

Citation for the whole project: Jepson Flora Project (eds.) 2018, Jepson eFlora,, accessed on July 19, 2018.

Geographic subdivisions for Lycopus asper:
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.