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Lappula squarrosa
TWO-ROWED STICKSEED

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key

Common Name: BORAGE or WATERLEAF FAMILY
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Genus: LappulaView DescriptionDichotomous Key

Habit: Annual, hairy, taprooted. Stem: +- erect; branches 0--many. Leaf: basal and cauline, sessile, entire. Inflorescence: raceme-like cymes, +- terminal; bracts leaf-like; pedicels erect in fruit. Flower: calyx +- deep-5-lobed, enlarged in fruit; corolla 5-lobed, funnel-shaped, appendages present; style entire. Fruit: nutlets generally 4, ovate, covered with +- long, barbed prickles, scar lateral.
Species In Genus: 12--14 species: northern hemisphere, especially Asia. Etymology: (Latin: little bur)

Lappula squarrosa (Retz.) Dumort.
WAIF
Stem: 1.5--8 dm. Leaf: 2--5 cm, linear to oblong. Inflorescence: pedicel 1--3 mm. Flower: calyx 2.5--3 mm in fruit, lobes wide-linear, spreading in fruit; corolla 2--3 mm diam, blue. Fruit: nutlets 3--4 mm, marginal prickles in 2(3) rows, bases not or +- widened, not fused. Chromosomes: 2n=48.
Ecology: Moist, disturbed areas; Elevation: < 200 m. Bioregional Distribution: SCo (Santa Monica), ScV (Lodi), expected MP; Distribution Outside California: to eastern Washington, southern Idaho, northern Utah, naturalized in northern North America, Canada; native to Eurasia. Flowering Time: May--Jul Note: Last collected 1941.
Synonyms: Lappula echinata Gilib.; Lappula fremontii (Torr.) Greene
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley
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Citation for this treatment: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley 2016. Lappula squarrosa, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=30244, accessed on February 09, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on February 09, 2016.


Geographic subdivisions for Lappula squarrosa:
SCo (Santa Monica), ScV (Lodi), expected MP;
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.