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Hydrophyllum tenuipes
SLENDERFOOT WATERLEAF, PACIFIC WATERLEAF

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key

Common Name: BORAGE or WATERLEAF FAMILY
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Genus: HydrophyllumView DescriptionDichotomous Key

Common Name: WATERLEAF
Habit: [Biennial or] perennial herb from rhizomes [taproots]; roots fleshy-fibrous and fibrous [or tuber-like]. Stem: suberect to erect, simple or branched, fleshy, or 0 and leaves from rhizomes. Leaf: simple, pinnate-[palmate-]lobed, or compound, basal or cauline, alternate, mottled white in shade; petiole widened, proximally purple, bases clasping, persistent, fleshy, juicy; leaflets toothed or lobed, hairy, generally paler abaxially, basal pair free, terminal +- united. Inflorescence: generally branched, generally head-like cymes; peduncles proximally purplish, in fruit erect or recurved; pedicels generally elongate, in fruit spreading or recurved. Flower: calyx bell-shaped, lobes generally equal, generally +- alike, linear to narrowly oblong or lanceolate, acute to obtuse, glabrous or hairy, generally ciliate, generally +- enlarging in fruit, sinus appendages 0 [present]; corolla lobed to middle, > calyx, bell-shaped, white, cream, +- green, purple, or blue, tube with linear scales at base forming channeled pollinator guide, lobes with nectary gland on midvein, hairy; stamens equal, exserted, filaments at mid-level generally hairy; ovary chamber 1, placentas parietal, style 1, exserted, glabrous, cleft < 1/4, stigmas 2, base persistent, disk proximal to ovary. Fruit: capsule, 3--5 mm, spheric; tip generally bristly, loosely enclosed by calyx. Seed: 1--4, oblong to spheric, brown or yellow- or red-brown, net-like, fleshy appendages 0. Chromosomes: 2n=18.
Species In Genus: 11 species: North America; some cultivated as ornamentals. Etymology: (Greek: water leaf) Note: Hydrophyllum capitatum var. alpinum raised to species rank, as Hydrophyllum alpestre.
eFlora Treatment Author: Genevieve K. Walden, Robert Patterson & Richard R. Halse

Hydrophyllum tenuipes A. Heller
NATIVE
Habit: Plant 20--80 cm; rhizome long. Stem: with reflexed bristles. Leaf: petiole 8--30 cm; blade 7--15 cm, wide-ovate to round; leaflets generally 3--5(9), lobed or not, coarse-serrate to cut, proximal pair(s) generally smaller, distinct, terminal merged or not (or appearing 3-lobed). Inflorescence: well above ground, >= subtending leaves; peduncle 2--15 cm, in fruit erect; pedicels 4--12 mm, in fruit to 15 mm, spreading or recurved. Flower: calyx lobes 4--7 mm; corolla 5--7 mm, lobes 3--4 mm, cream, +- green [purple or blue]; anthers 1--2 mm; style 9--14 mm. Seed: 1(-3), 2.5--3.5 mm, yellow- or red-brown.
Ecology: Moist, shaded, wooded slopes, streambanks; Elevation: < 1750 m. Bioregional Distribution: NCo, KR, NCoRO; Distribution Outside California: to British Columbia. Flowering Time: Apr--Jul
Synonyms: Hydrophyllum viridulum G.N. Jones, Hydrophyllum tenuipes var. viride Jeps.
eFlora Treatment Author: Genevieve K. Walden, Robert Patterson & Richard R. Halse
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botanical illustration including Hydrophyllum tenuipes

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Citation for this treatment: Genevieve K. Walden, Robert Patterson & Richard R. Halse 2016. Hydrophyllum tenuipes, Revision 2, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=28635, accessed on May 27, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on May 27, 2016.


Geographic subdivisions for Hydrophyllum tenuipes:
NCo, KR, NCoRO;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.