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Howellia aquatilis

Higher Taxonomy
Family: CampanulaceaeView DescriptionDichotomous Key

Habit: Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, +- fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or +- fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1--3, placentas axile or parietal, ovules many, style generally 1, 2--5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
Genera In Family: +- 90 genera, +- 2500 species: worldwide. Note: Some cultivated for ornamental (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus.
eFlora Treatment Author: Nancy R. Morin, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: HowelliaView Description 

Species In Genus: 1 sp. Etymology: (Thomas (1842--1912) and Joseph (1830--1912) Howell, brothers and botanists in Pacific Northwest)

Howellia aquatilis A. Gray
Habit: Annual, aquatic, rooted, glabrous. Stem: submersed, ascending to erect, or floating; 10--60 cm. Leaf: alternate (sometimes +- opposite or whorled), cauline, linear, entire or minutely toothed, sessile. Inflorescence: flowers 1 in leaf or bract axils, scattered; pedicels stout, 1--4(8) mm; bract 1 per flower. Flower: inverted; sepals 1/6--2/5 ovary, narrowly triangular; corolla white, 0 in lower flowers or not, tube 2--2.7 mm, narrowly cylindric, split adaxially +- to base, +- 2-lipped, 2 adaxial lobes +- 1--1.3 mm, narrowly strap-shaped, spreading, 3 abaxial lobes +- 1--1.3 mm; stamens fused into tube, anthers 0.2--0.9 mm, 2 shorter anthers minutely appendaged; ovary inferior, 5--13 mm, ellipsoid, stigma head-like. Fruit: 5--13 mm, 1--2 mm diam, ellipsoid, 1-chambered, depressed-conic at top, open on sides by slits or irregular tears. Seed: 2--4 mm, cylindric, shiny, chestnut-brown.
Ecology: Seasonal ponds; Elevation: 1100--1500 m. Bioregional Distribution: NCoRH; Distribution Outside California: to Washington, Montana. Flowering Time: Jun--Aug
eFlora Treatment Author: Nancy R. Morin
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Listed on CNPS Rare Plant Inventory

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botanical illustration including Howellia aquatilis


Citation for this treatment: Nancy R. Morin 2016. Howellia aquatilis, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 05, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on May 05, 2016.

Howellia aquatilis
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© 2005 Dieter Wilken

More photos of Howellia aquatilis in CalPhotos

Geographic subdivisions for Howellia aquatilis:
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.