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Hackelia velutina
VELVETY STICKSEED

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key

Common Name: BORAGE or WATERLEAF FAMILY
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Genus: HackeliaView DescriptionDichotomous Key

Common Name: STICKSEED
Habit: Perennial herb (biennial); hairs appressed to spreading; caudex generally branched in age, often +- woody, taprooted. Stem: ascending or erect. Leaf: lowest petioles generally +- = blades, +- winged, others 0. Inflorescence: coiled cymes, generally > 3, generally terminal and axillary, +- bracted; pedicel in fruit elongated, recurved to reflexed. Flower: calyx deep-5-lobed; corolla rotate-salverform, generally white with yellow patch adaxially, lobes appendaged near base. Fruit: nutlets erect, > style, attachment scar lateral-medial, generally with barb-tipped prickles abaxially and on margin.
Species In Genus: 40 species: generally western North America, southeastern Asia. Etymology: (J. Hackel, Czech botanist, 1783--1869) Note: Values for corolla limb diam take into account shrinkage during flower period. Difficult, study needed, especially in northern California, southeastern Asia; sometimes merged with Lappula.
Unabridged Note: In North America filling old-world role of flat-flowered forget-me-nots, Myosotis, Lepechiniella, etc, for pollinators.
eFlora Treatment Author: Ronald B. Kelley & Robert L. Carr

Hackelia velutina (Piper) I.M. Johnst.
NATIVE
Stem: 4--8 dm; hairs generally spreading, +- dense below middle, +- dense to +- sparse above. Leaf: velvety-hairy; basal generally 5--17 cm, 0.5--2 cm wide, narrow-elliptic to oblanceolate; lower cauline reduced, ovate to narrow-lanceolate, +- clasping upward. Inflorescence: +- dense, branches several, few-flowered; pedicel 6--20 mm in fruit. Flower: calyx 2--3 mm; corolla tube generally >> calyx, generally +- purple, throat +- closed, limb 12--20 mm diam, generally blue to +- purple (pink), appendages long-exserted, much longer than wide, recurved, 2-pronged at tip, white; anthers hidden. Fruit: nutlets 4.5--6.5 mm, dull, roughened, abaxial prickles many, evenly distributed, +- = marginal.
Ecology: Dry, open slopes, forest clearings, roadsides; Elevation: 1350--2750 m. Bioregional Distribution: SNH; Distribution Outside California: western Nevada. Flowering Time: Jun--Aug
Synonyms: Hackelia longituba I.M. Johnst.
Unabridged Note: Though corolla distinctive, mistaken for Hackelia nervosa, Hackelia mundula: the unique long-exserted recurved corolla appendages and overall velvety hairs distinguish it from Hackelia nervosa; the exserted corolla tube, white corolla appendages, and dull, roughened nutlets separate it from Hackelia mundula. A CalPhoto image taken in southern SN shows a pink flowered pl, no doubt indicating introgression with Hackelia mundula. The suspected occurrence of Hackelia velutina in CaRH based on multiple CalPhoto images in the Lassen Volcanic National Park area, though it is uncertain whether voucher specimens exist.
eFlora Treatment Author: Ronald B. Kelley & Robert L. Carr
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Citation for this treatment: Ronald B. Kelley & Robert L. Carr 2016. Hackelia velutina, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=27544, accessed on December 09, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on December 09, 2016.


Hackelia velutina
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© 2015 Neal Kramer
Hackelia velutina
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© 2009 Keir Morse
Hackelia velutina
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© 2015 Barry Rice
Hackelia velutina
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© 2012 Gary A. Monroe
Hackelia velutina
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© 2009 Barry Breckling
Hackelia velutina
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© 2015 Neal Kramer

More photos of Hackelia velutina in CalPhotos


Geographic subdivisions for Hackelia velutina:
SNH;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.