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Hackelia nervosa
SIERRAN STICKSEED

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key

Common Name: BORAGE or WATERLEAF FAMILY
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Genus: HackeliaView DescriptionDichotomous Key

Common Name: STICKSEED
Habit: Perennial herb (biennial); hairs appressed to spreading; caudex generally branched in age, often +- woody, taprooted. Stem: ascending or erect. Leaf: lowest petioles generally +- = blades, +- winged, others 0. Inflorescence: coiled cymes, generally > 3, generally terminal and axillary, +- bracted; pedicel in fruit elongated, recurved to reflexed. Flower: calyx deep-5-lobed; corolla rotate-salverform, generally white with yellow patch adaxially, lobes appendaged near base. Fruit: nutlets erect, > style, attachment scar lateral-medial, generally with barb-tipped prickles abaxially and on margin.
Species In Genus: 40 species: generally western North America, southeastern Asia. Etymology: (J. Hackel, Czech botanist, 1783--1869) Note: Values for corolla limb diam take into account shrinkage during flower period. Difficult, study needed, especially in northern California, southeastern Asia; sometimes merged with Lappula.
Unabridged Note: In North America filling old-world role of flat-flowered forget-me-nots, Myosotis, Lepechiniella, etc, for pollinators.
eFlora Treatment Author: Ronald B. Kelley & Robert L. Carr

Hackelia nervosa (Kellogg) I.M. Johnst.
NATIVE
Stem: 4--7 dm; +- glabrous above, hairs generally sparse below, often +- spreading. Leaf: sparse-rough-hairy; basal 3--12 cm, 0.6--2.5 cm wide, +- oblong to oblanceolate; lower cauline generally smaller, generally sessile, ovate to lanceolate upward; uppermost +- clasping. Inflorescence: open, branches few--several; pedicel 6--12 mm in fruit. Flower: calyx 1.5--3.5+ mm; corolla tube > calyx, +- white, throat open, limb (8)10--18 mm diam, +- blue, appendages generally as long as wide, indented at tip; anthers visible at throat. Fruit: nutlets 4--6 mm, dull, roughened, abaxial prickles many, evenly distributed, +- = marginal.
Ecology: Moist open slopes, openings in forest; Elevation: 1800--2930 m. Bioregional Distribution: s CaRH, n&c SNH; Distribution Outside California: western Nevada. Flowering Time: May--Aug
Unabridged Note: Much larger corollas, nutlet differences relative to sympatric Hackelia micrantha.
eFlora Treatment Author: Ronald B. Kelley & Robert L. Carr
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Citation for this treatment: Ronald B. Kelley & Robert L. Carr 2016. Hackelia nervosa, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=27537, accessed on July 23, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on July 23, 2016.


Hackelia nervosa
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© 2003 Christopher L. Christie
Hackelia nervosa
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© 2009 Barry Breckling
Hackelia nervosa
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© 2009 Barry Breckling
Hackelia nervosa
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© 1995 Gary A. Monroe
Hackelia nervosa
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© 2015 Barry Rice
Hackelia nervosa
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© 2003 Christopher L. Christie

More photos of Hackelia nervosa in CalPhotos


Geographic subdivisions for Hackelia nervosa:
s CaRH, n&c SNH;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.