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Vascular Plants of California
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Gratiola heterosepala
BOGGS LAKE HEDGE-HYSSOP


Higher Taxonomy
Family: PlantaginaceaeView DescriptionDichotomous Key
Common Name: PLANTAIN FAMILY
Habit: Annual to shrub, some aquatic. Leaf: basal or cauline, alternate or opposite (whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0. Inflorescence: raceme, spike, or flowers axillary in 1--few-flowered clusters; flowers few to many, each subtended by 1 bract. Flower: unisexual or bisexual, radial or bilateral; sepals 4--5, generally fused at base; corolla 4--5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1--2, anthers opening by 2 slits; ovary superior, [1]2--4-chambered, style 1, stigma lobes 0 or 2. Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
Genera In Family: 110 genera, +- 2000 species: worldwide, especially temperate. Note: Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic California genera of Scrophulariaceae (except Buddleja, Diplacus, Erythranthe, Limosella, Mimetanthe, Mimulus, Myoporum, Scrophularia, Verbascum). California Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated.
eFlora Treatment Author: Margriet Wetherwax, except as noted
Scientific Editor: Robert Patterson, Bruce G. Baldwin.
Genus: GratiolaView DescriptionDichotomous Key


Common Name: HEDGE-HYSSOP
Habit: Annual; herbage glabrous proximally, glandular-puberulent distally; roots fibrous. Stem: erect to decumbent, < 30 cm, rounded to +- 4-angled. Leaf: cauline, opposite, sessile or +- clasping, subentire to dentate, palmately veined. Inflorescence: flowers 1 per leaf axil, pedicelled, bractlets present or 0. Flower: sepals 5, generally free; corolla 5-lobed, tube 4-angled, > lobes, +- purple-lined, upper lip 2-lobed or notched, lower 3-lobed; fertile stamens 2, anther sacs of each stamen separated, parallel; ovary subtended by nectary, stigma weakly 2-lobed. Fruit: septicidal and loculicidal, 4-valved, 3--6 mm, ovoid to spheric. Seed: many, 0.5--0.9 mm, coat net-like, +- brown.
Species In Genus: +- 25 species: temperate, tropical mountains, North America, Europe, Asia, Australia, South America. Etymology: (Latin: grace or favor, from supposed medicinal quality)
eFlora Treatment Author: Dwayne Estes
Reference: Estes & Small 2008 Syst Bot 33:176--182
Gratiola heterosepala H. Mason & Bacig.
NATIVE
Stem: erect, generally simple, 2--10 cm, with 4--7 leafy nodes. Leaf: 2--20 mm, proximal lance-linear, oblong to obovate distally on stem, distal +- clasping, tip rounded to notched, margin entire. Inflorescence: pedicels 5--25 mm, slender, erect; bractlets 0. Flower: sepals unequal, unequally fused at base, 3.5--5 mm, to 6 mm in fruit, elliptic-oblanceolate to obovate, tip rounded to notched; corolla 5--8 mm, generally yellow, except lower 3 lobes white. Fruit: 3.5--5 mm, ovoid, valve tips blunt. Seed: 0.5--0.7 mm, 0.2--0.3 mm wide.
Ecology: Shallow water, margins of vernal pools; Elevation: < 1600 m (2400 m in Wrn). Bioregional Distribution: NCoRI, CaR, n&c SNF, GV, MP; Distribution Outside California: to south-central Oregon (Lake Co.). Flowering Time: Apr--Sep
Jepson eFlora Author: Dwayne Estes
Reference: Estes & Small 2008 Syst Bot 33:176--182
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Listed on CNPS Rare Plant Inventory

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Botanical illustration including Gratiola heterosepala

botanical illustration including Gratiola heterosepala

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Citation for this treatment: Dwayne Estes 2012, Gratiola heterosepala, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=27295, accessed on November 12, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora, http://ucjeps.berkeley.edu/eflora/, accessed on November 12, 2019.

Gratiola heterosepala
click for enlargement
© 2004 Carol W. Witham
Gratiola heterosepala
click for enlargement
© 2004 Carol W. Witham
Gratiola heterosepala
click for enlargement
© 2005 Robert E. Preston, Ph.D.

More photos of Gratiola heterosepala in CalPhotos



Geographic subdivisions for Gratiola heterosepala:
NCoRI, CaR, n&c SNF, GV, MP
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

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Data provided by the participants of the  Consortium of California Herbaria.
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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.