Jepson eFlora: Taxon page
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Gratiola heterosepala
BOGGS LAKE HEDGE-HYSSOP

Higher Taxonomy
Family: PlantaginaceaeView DescriptionDichotomous Key

Common Name: PLANTAIN FAMILY
Habit: Annual to shrub, some aquatic. Leaf: basal or cauline, alternate or opposite (whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0. Inflorescence: raceme, spike, or flowers axillary in 1--few-flowered clusters; flowers few to many, each subtended by 1 bract. Flower: unisexual or bisexual, radial or bilateral; sepals 4--5, generally fused at base; corolla 4--5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1--2, anthers opening by 2 slits; ovary superior, [1]2--4-chambered, style 1, stigma lobes 0 or 2. Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
Genera In Family: 110 genera, +- 2000 species: worldwide, especially temperate. Note: Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic California genera of Scrophulariaceae (except Buddleja, Limosella, Mimulus, Myoporum, Scrophularia, Verbascum). California Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated.
eFlora Treatment Author: Margriet Wetherwax, except as noted
Scientific Editor: Robert Patterson, Bruce G. Baldwin.
Genus: GratiolaView DescriptionDichotomous Key

Common Name: HEDGE-HYSSOP
Habit: Annual; herbage glabrous proximally, glandular-puberulent distally; roots fibrous. Stem: erect to decumbent, < 30 cm, rounded to +- 4-angled. Leaf: cauline, opposite, sessile or +- clasping, subentire to dentate, palmately veined. Inflorescence: flowers 1 per leaf axil, pedicelled, bractlets present or 0. Flower: sepals 5, generally free; corolla 5-lobed, tube 4-angled, > lobes, +- purple-lined, upper lip 2-lobed or notched, lower 3-lobed; fertile stamens 2, anther sacs of each stamen separated, parallel; ovary subtended by nectary, stigma weakly 2-lobed. Fruit: septicidal and loculicidal, 4-valved, 3--6 mm, ovoid to spheric. Seed: many, 0.5--0.9 mm, coat net-like, +- brown.
Species In Genus: +- 25 species: temperate, tropical mountains, North America, Europe, Asia, Australia, South America. Etymology: (Latin: grace or favor, from supposed medicinal quality)
eFlora Treatment Author: Dwayne Estes

Gratiola heterosepala H. Mason & Bacig.
NATIVE
Stem: erect, generally simple, 2--10 cm, with 4--7 leafy nodes. Leaf: 2--20 mm, proximal lance-linear, oblong to obovate distally on stem, distal +- clasping, tip rounded to notched, margin entire. Inflorescence: pedicels 5--25 mm, slender, erect; bractlets 0. Flower: sepals unequal, unequally fused at base, 3.5--5 mm, to 6 mm in fruit, elliptic-oblanceolate to obovate, tip rounded to notched; corolla 5--8 mm, generally yellow, except lower 3 lobes white. Fruit: 3.5--5 mm, ovoid, valve tips blunt. Seed: 0.5--0.7 mm, 0.2--0.3 mm wide.
Ecology: Shallow water, margins of vernal pools; Elevation: < 1600 m (2400 m in Wrn). Bioregional Distribution: NCoRI, CaR, n&c SNF, GV, MP; Distribution Outside California: to south-central Oregon (Lake Co.). Flowering Time: Apr--Sep
eFlora Treatment Author: Dwayne Estes
Jepson Online Interchange
Listed on CNPS Rare Plant Inventory

Previous taxon: Gratiola ebracteata
Next taxon: Gratiola neglecta

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botanical illustration including Gratiola heterosepala

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Citation for this treatment: Dwayne Estes 2016. Gratiola heterosepala, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=27295, accessed on April 29, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on April 29, 2016.


Gratiola heterosepala
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© 2004 Carol W. Witham
Gratiola heterosepala
click for enlargement
© 2004 Carol W. Witham
Gratiola heterosepala
click for enlargement
© 2005 Robert E. Preston, Ph.D.

More photos of Gratiola heterosepala in CalPhotos


Geographic subdivisions for Gratiola heterosepala:
NCoRI, CaR, n&c SNF, GV, MP;
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.