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Vascular Plants of California
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Gratiola ebracteata
BRACTLESS HEDGE-HYSSOP


Higher Taxonomy
Family: PlantaginaceaeView DescriptionDichotomous Key
Common Name: PLANTAIN FAMILY
Habit: Annual to shrub, some aquatic. Leaf: basal or cauline, alternate or opposite (whorled), simple, entire to dentate or lobed, venation generally pinnate; stipules 0. Inflorescence: raceme, spike, or flowers axillary in 1--few-flowered clusters; flowers few to many, each subtended by 1 bract. Flower: unisexual or bisexual, radial or bilateral; sepals 4--5, generally fused at base; corolla 4--5-lobed, scarious or not, persistent or not, generally 2-lipped, upper lip generally 2-lobed, lower generally 3-lobed, spur present or not, tube sac-like at base or not; stamens 2 or 4, alternate corolla lobes, epipetalous, staminode 0 or 1--2, anthers opening by 2 slits; ovary superior, [1]2--4-chambered, style 1, stigma lobes 0 or 2. Fruit: generally a capsule, septicidal, loculicidal, circumscissile, or dehiscing by terminal slits or pores.
Genera In Family: 110 genera, +- 2000 species: worldwide, especially temperate. Note: Veronicaceae sensu Olmstead et al. Recently treated to include Callitrichaceae, Hippuridaceae, and most non-parasitic California genera of Scrophulariaceae (except Buddleja, Diplacus, Erythranthe, Limosella, Mimetanthe, Mimulus, Myoporum, Scrophularia, Verbascum). California Maurandya moved to Holmgrenanthe and Maurandella. Limnophila ×ludoviciana Thieret an occasional agricultural weed in rice fields. Hebe ×franciscana (Eastw.) Souster, Hebe speciosa (R. Cunn.) Andersen only cultivated.
eFlora Treatment Author: Margriet Wetherwax, except as noted
Scientific Editor: Robert Patterson, Bruce G. Baldwin.
Genus: GratiolaView DescriptionDichotomous Key


Common Name: HEDGE-HYSSOP
Habit: Annual; herbage glabrous proximally, glandular-puberulent distally; roots fibrous. Stem: erect to decumbent, < 30 cm, rounded to +- 4-angled. Leaf: cauline, opposite, sessile or +- clasping, subentire to dentate, palmately veined. Inflorescence: flowers 1 per leaf axil, pedicelled, bractlets present or 0. Flower: sepals 5, generally free; corolla 5-lobed, tube 4-angled, > lobes, +- purple-lined, upper lip 2-lobed or notched, lower 3-lobed; fertile stamens 2, anther sacs of each stamen separated, parallel; ovary subtended by nectary, stigma weakly 2-lobed. Fruit: septicidal and loculicidal, 4-valved, 3--6 mm, ovoid to spheric. Seed: many, 0.5--0.9 mm, coat net-like, +- brown.
Species In Genus: +- 25 species: temperate, tropical mountains, North America, Europe, Asia, Australia, South America. Etymology: (Latin: grace or favor, from supposed medicinal quality)
eFlora Treatment Author: Dwayne Estes
Reference: Estes & Small 2008 Syst Bot 33:176--182
Gratiola ebracteata A. DC.
NATIVE
Stem: simple to branched, 5--20 cm, with 6--12 leafy nodes. Leaf: 7--25 mm, lance-ovate to lance-linear, acuminate, generally subentire. Inflorescence: pedicels 5--25 mm, stout, erect; bractlets 0. Flower: sepals free, equal, 4--7 mm, 6--15 mm in fruit, lanceolate, acuminate; corolla 5--8 mm, throat +- yellow, limb white +- pink-tinged. Fruit: 3.5--6 mm, spheric, 4-angled in ×-section, valve tips abruptly pointed. Seed: 0.6--0.9 mm, 0.3--0.4 mm wide.
Ecology: Wet, muddy places; Elevation: < 2400 m. Bioregional Distribution: n&c CA-FP (exc NCoRH), MP; Distribution Outside California: to British Columbia, Montana. Flowering Time: Apr--Jun
Jepson eFlora Author: Dwayne Estes
Reference: Estes & Small 2008 Syst Bot 33:176--182
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Botanical illustration including Gratiola ebracteata

botanical illustration including Gratiola ebracteata

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Citation for this treatment: Dwayne Estes 2012, Gratiola ebracteata, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=27292, accessed on May 22, 2019.

Citation for the whole project: Jepson Flora Project (eds.) 2019, Jepson eFlora, http://ucjeps.berkeley.edu/eflora/, accessed on May 22, 2019.

Gratiola ebracteata
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© 2009 Aaron Schusteff
Gratiola ebracteata
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© 2017 Chris Winchell
Gratiola ebracteata
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© 2010 Barry Breckling
Gratiola ebracteata
click for enlargement
© 2010 Barry Breckling
Gratiola ebracteata
click for enlargement
© 2017 Chris Winchell
Gratiola ebracteata
click for enlargement
© 2004 Carol W. Witham

More photos of Gratiola ebracteata in CalPhotos



Geographic subdivisions for Gratiola ebracteata:
n&c CA-FP (exc NCoRH), MP
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

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Data provided by the participants of the  Consortium of California Herbaria.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.