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    Vascular Plants of California
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Ephedra aspera

Higher Taxonomy
Family: EphedraceaeView Description 

Habit: Shrub; generally dioecious; taproot deep. Stem: erect, prostrate, occasionally climbing, jointed, much branched, green in youth; branchlets opposite or whorled, cylindric, longitudinally grooved; pith brown or white. Leaf: simple, 2 (each pair at right angles to adjacent pairs) or 3 per node, scale-like, +- fused at base into membranous sheath, generally ephemeral; resin canals 0. Cone: axillary (terminal), sessile or not, 1--many per node, ovoid or ellipsoid. Pollen Cone: bracts 2--3 per node, proximal sterile, distal subtending 2 basally fused bractlets subtending a stalk with 2--10(15) sessile to stalked, anther-like parts with 2 apical pores. Seed Cone: stalk 1--11(50) mm, generally straight, naked or with reduced basal bracts; distal bracts in (2)4--10 pairs or whorls, free or fused, brown or green, scarious or fleshy, subtending 2 fused leathery bractlets enclosing 1 ovule. Seed: 1--3(4) per cone (in ×-section, +- round if 1, +- semicircular if 2, +- triangular if 3, etc.), +- spheric to ellipsoid or lance-ovoid, beaked or not.
Genera In Family: 1 genus, +- 50 species: arid, semi-arid (to tropical) regions of northern Africa, Asia, Europe, North America, South America.
eFlora Treatment Author: Stefanie M. Ickert-Bond
Scientific Editor: Thomas J. Rosatti.
Genus: EphedraView DescriptionDichotomous Key

Stem: spreading to generally erect, 0.3--2(4.5) m; uppermost bud generally conic, generally not thorn-like.
Etymology: (Greek: sitting upon, from jointed stems and/or ancient name for Equisetum, also with jointed stems)

Ephedra aspera S. Watson
Stem: 0.3--2 m; twig +- green, aging yellow, glaucous or not; pith dark brown. Leaf: 2 per node, 1--3.5 mm, generally persistent; sheath splitting, base in age swollen. Pollen Cone: 1--many per node, 2.5--7 mm; bracts 2 per node. Seed Cone: 1--few per node, 4.5--8(10) mm, ovoid, stalk <= 2 mm, basal bracts reduced; distal bracts 2 per node, free, ovate, red-brown, margins membranous, stalk-like base 0. Seed: generally 1, 4--7(8) mm, generally < 2 × as long as wide, smooth to +- roughened. Chromosomes: 2n=14,28.
Ecology: Creosote-bush scrub, Joshua-tree woodland; Elevation: 50--2133 m. Bioregional Distribution: SnBr, SnJt, SNE, D; Distribution Outside California: to Texas, Mexico. Note: Cones present Mar--Apr.
Synonyms: Ephedra clokeyi H.C. Cutler; Ephedra fasciculata A. Nelson; Ephedra fasciculata var. clokeyi (H.C. Cutler) Clokey; Ephedra fasciculata var. fasciculata; Ephedra nevadensis S. Watson var. aspera (Engelm. ex S. Watson) L.D. Benson; Ephedra peninsularis I.M. Johnst.; Ephedra reedii Cory;
Jepson eFlora Author: Stefanie M. Ickert-Bond
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botanical illustration including Ephedra aspera


Citation for this treatment: Stefanie M. Ickert-Bond 2012, Ephedra aspera, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on November 17, 2018.

Citation for the whole project: Jepson Flora Project (eds.) 2018, Jepson eFlora,, accessed on November 17, 2018.

Geographic subdivisions for Ephedra aspera:
SnBr, SnJt, SNE, D;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time.