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Cuscuta howelliana
BOGGS LAKE DODDER

Higher Taxonomy
Family: ConvolvulaceaeView DescriptionDichotomous Key

Common Name: MORNING-GLORY FAMILY
Habit: Annual, perennial herb, subshrub, generally twining or trailing. Leaf: 0 or alternate. Inflorescence: cyme or flowers 1 in axils; bracts subtending flowers 0 or 2. Flower: bisexual, radial; sepals (4)5, +- free, overlapping, persistent, often unequal; corolla generally showy, generally bell-shaped, +- shallowly 5-lobed, generally pleated and twisted in bud; stamens 5, epipetalous; pistil 1, ovary superior, chambers generally 2, each generally 2-ovuled, styles 1--2. Fruit: generally capsule. Seed: 1--4(6).
Genera In Family: 55--60 genera, 1600--1700 species: warm temperate to tropics; some cultivated for food or as ornamental (Ipomoea). Note: Monophyletic only if Cuscutaceae included, as treated here. Ipomoea cairica (L.) Sweet, Ipomoea hederacea Jacq. [Ipomoea nil L., misappl.], Ipomoea indica (Burm.) Merr. (including Ipomoea mutabilis Ker Gawl.), Ipomoea purpurea (L.) Roth, Ipomoea triloba L., all included in TJM (1993), not naturalized.
eFlora Treatment Author: Robert E. Preston, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: CuscutaView DescriptionDichotomous Key

Common Name: DODDER
Habit: Vine, annual (perennial herb if on perennial host), not in contact with ground, attached to, holoparasitic on host by many small, specialized roots (haustoria) along stem, generally glabrous. Stem: thread-like, +- green, yellow, orange, or +- red. Leaf: 0 or scale-like, alternate, +- 2 mm. Inflorescence: generally cyme, head- to panicle-like (flowers 1), subtended by 0--3 bracts. Flower: bisexual, radial, parts generally in 4s or 5s; calyx generally divided 2/5--3/5, persistent generally +- cream-white; corolla generally +- white, persistent (withered in fruit) or not, tube cup-shaped to cylindric, bulged or horizontally ridged below lobes or generally not, generally with scales subtending stamens, lobes alternate stamens, erect to reflexed; ovary superior, chambers 2, each 2-ovuled, styles 2, generally free, persistent, stigmas 2, generally spheric, persistent. Fruit: capsule, generally indehiscent to irregularly dehiscent (or circumscissile near base), spheric to ovoid, depressed or not, thickened and/or raised around generally inconspicuous opening between styles or not. Seed: 1--4; coat papillate when hydrated, honeycombed when dry, (rarely neither, with cells +- rectangular, in +- jigsaw-puzzle-like arrangement); embryo generally slender, 1--3-coiled.
Species In Genus: +- 180 species: cosmopolitan, especially warmer regions of western hemisphere and Polynesia. Etymology: (Aramaic, Hebrew; from the verb K-S-Y (Kaph, Shin, Yodh), to cover, from habit) Note: By persistent, withered corolla, fruit may be "capped" (corolla on top of fruit), "surrounded" (fruit at least in part visible, corolla +- loosely around fruit), or "enclosed" (fruit not visible, corolla +- tightly around fruit). Cuscuta pentagona Engelm. excluded.
eFlora Treatment Author: Mihai Costea & Saša Stefanović

Cuscuta howelliana P. Rubtzov
NATIVE
Inflorescence: in inflorescence of host, head-like, flowers 3--30; pedicels 0--0.6 mm. Flower: 3--4.5 mm, membranous, papillate, parts in 4s(5s); calyx >= corolla tube, bell-shaped, divided 1/2--2/3, finely veined, shiny, lobes triangular-ovate, bases not overlapped, margins entire, tip acuminate to long-acuminate, recurved; corolla tube 1.5--2.2 mm, narrowly bell- to urn-shaped, scales +- 1/2 corolla tube, oblong-ovate, rounded, uniformly densely fringed, lobes +- erect to spreading, generally = tube, triangular-ovate, margins entire, tip acute to long-acuminate, recurved; filaments 0.1--0.4 mm, anthers included, 0.4--0.7 mm, elliptic; styles 0.4--1.1 mm, 1/4 to generally = ovary. Fruit: 1.2--1.5 mm, 0.8--1.2 mm wide, spheric to spheric-depressed, not thickened or raised around opening between styles, not translucent, enclosed by corolla, capped by it in age. Seed: 1--4, 0.9--1.4 mm, 0.8--1.3 mm wide, +- round to widely elliptic.
Ecology: Margins of vernal pools generally on Eryngium, Navarretia, Polygonum polygaloides subsp. kelloggii, Epilobium campestre; Elevation: 140--1650 m. Bioregional Distribution: KR/CaRH, NCoRI, CaRF, n SNF, GV, MP. Flowering Time: Aug--Sep
eFlora Treatment Author: Mihai Costea & Saša Stefanović
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Next taxon: Cuscuta indecora var. indecora

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Citation for this treatment: Mihai Costea & Saša Stefanović 2016. Cuscuta howelliana, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=21447, accessed on May 05, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on May 05, 2016.


Cuscuta howelliana
click for enlargement
© 2004 Carol W. Witham
Cuscuta howelliana
click for enlargement
© 2004 Carol W. Witham
Cuscuta howelliana
click for enlargement
© 2004 Carol W. Witham
Cuscuta howelliana
click for enlargement
© 2006 George W. Hartwell
Cuscuta howelliana
click for enlargement
© 2004 Carol W. Witham
Cuscuta howelliana
click for enlargement
© 2004 Carol W. Witham

More photos of Cuscuta howelliana in CalPhotos


Geographic subdivisions for Cuscuta howelliana:
KR/CaRH, NCoRI, CaRF, n SNF, GV, MP.
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.