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Cardamine angulata

Higher Taxonomy
Family: Brassicaceae (Cruciferae)View DescriptionDichotomous Key

Common Name: MUSTARD FAMILY
Habit: Annual to shrub; sap pungent, watery. Leaf: generally simple, alternate; generally both basal, cauline; stipules 0. Inflorescence: generally raceme, generally not bracted. Flower: bisexual, generally radial; sepals 4, generally free; petals (0)4, forming a cross, generally white or yellow to purple; stamens generally 6 (2 or 4), 4 long, 2 short (3 pairs of unequal length); ovary 1, superior, generally 2-chambered with septum connecting 2 parietal placentas; style 1, stigma entire or 2-lobed. Fruit: capsule, generally 2-valved, "silique" (length >= 3 × width) or "silicle" (length < 3 × width), dehiscent by 2 valves or indehiscent, cylindric or flat parallel or perpendicular to septum, segmented or not. Seed: 1--many, in 1 or 2 rows per chamber, winged or wingless; embryo strongly curved.
Genera In Family: +- 330 genera, 3780 species: worldwide, especially temperate. Note: Highest diversity in Mediterranean area, mountains of southwestern Asia, adjacent central Asia, western North America; some Brassica species are oil or vegetable crops; Arabidopsis thaliana used in experimental molecular biology; many species are ornamentals, weeds. Aurinia saxatilis (L.) Desvaux in cultivation only. Aubrieta occasional waif in central NCoR, Carrichtera annua (L.) DC. in SCo, Iberis sempervirens L., Iberis umbellata L. in PR, Teesdalia coronopifolia (Bergeret) Thell., Teesdalia nudicaulis (L.) W.T. Aiton in southern NCoRO, CCo. Cardaria, Coronopus moved to Lepidium; Caulostramina to Hesperidanthus; Guillenia to Caulanthus; Heterodraba to Athysanus; California taxa of Lesquerella to Physaria; Malcolmia africana to Strigosella.
eFlora Treatment Author: Ihsan A. Al-Shehbaz, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.
Genus: CardamineView DescriptionDichotomous Key

Common Name: BITTER-CRESS
Habit: Annual to perennial herb, from taproots, fibrous roots, or tuber-like rhizomes; hairs 0 or simple. Leaf: alternate, opposite, or whorled; entire to palmately, pinnately lobed, or compound; cauline leaves petioled or 0, not lobed at base [lobed]. Inflorescence: elongated, bracts generally 0. Flower: sepals erect (spreading), bases sac-like or not; petals white, pink, purple, or violet. Fruit: silique, linear, flat parallel to septum, dehiscent, unsegmented; valves generally coiling when dehiscent; placental margins flattened. Seed: (4)10--80, in 1 row, wingless.
Species In Genus: +- 200 species: temperate, worldwide. Etymology: (Greek: for cress) Note: Some North American species (e.g., Cardamine californica, Cardamine nuttallii, Cardamine pachystigma) highly variable, more study needed; species treated conservatively here. Cardamine flexuosa With. a waif in gardens, nurseries.
Unabridged Note: Some North American species (e.g., Cardamine californica, Cardamine nuttallii, Cardamine pachystigma) highly variable and numerous minor variants were recognized as varieties. C ritical studies of these complexes reveal that only a fraction of the overall continuous variation was formally recognized. Without detailed molecular and cytological studies, a broader sp. concept is adopted herein.

Cardamine angulata Hook.
NATIVE
Habit: Perennial herb; rhizome slender, <= 2 mm wide, cylindric. Stem: simple, (1.5)2.5--8.5(10) dm; hairs at base sparse to dense. Leaf: leaflets of rhizome leaves 3(5), petiole (2)4--12(14) cm; cauline leaves (3)4--8, leaflets 3(5), ovate to lanceolate, margin dentate or entire. Flower: sepals 2.5--4 mm; petals 8--15 mm, 4--8 mm wide, white to +- pink. Fruit: erect to ascending, 1.5--3.2 cm, 1.4--2 mm wide; style (0.5)1--4 mm; pedicel ascending to spreading, (0.9)1.2--2.5 cm. Seed: 6--16(24), 1.8--2.3 mm, oblong. Chromosomes: 2n=40.
Ecology: Shady thickets, streambanks, forest; Elevation: < 900 m. Bioregional Distribution: NW; Distribution Outside California: to Alaska. Flowering Time: Apr--Jun
eFlora Treatment Author: Ihsan A. Al-Shehbaz
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Listed on CNPS Rare Plant Inventory

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Citation for this treatment: Ihsan A. Al-Shehbaz 2016. Cardamine angulata, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=17120, accessed on July 25, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on July 25, 2016.


Cardamine angulata
click for enlargement
© 2016 Dana York
Cardamine angulata
click for enlargement
© 2015 Dana York
Cardamine angulata
click for enlargement
© 2015 Dana York
Cardamine angulata
click for enlargement
© 2015 Dana York
Cardamine angulata
click for enlargement
© 2015 Dana York
Cardamine angulata
click for enlargement
© 2016 Dana York

More photos of Cardamine angulata in CalPhotos


Geographic subdivisions for Cardamine angulata:
NW;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.