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Botrychium pinnatum
NORTHWESTERN MOONWORT

Higher Taxonomy
Family: OphioglossaceaeView DescriptionDichotomous Key

Common Name: ADDER'S-TONGUE FAMILY
Habit: Perennial herb, small, fleshy, generally glabrous; caudex generally underground, unbranched; roots glabrous with bulblets or plantlets or not. Leaf: generally 1 per caudex per year, divided into 2 facing parts with a common stalk,(0)1 sterile and 1(2) fertile (fertile occasionally aborted); sterile photosynthetic part (trophophore) separated from spore-bearing part (sporophore) at to well above ground level; trophophore simple to compound, veins free and forked or netted with included veinlets; sporophore simple to compound, or 0 in young pls. Sporangia: dehiscent into 2 valves, +- 1 mm wide, thick-walled.
Genera In Family: 10 genera, 80--100 species: +- worldwide, generally rare or overlooked.
Unabridged Note: Distantly related to most (leptosporangiate) ferns. Haploid (gametophyte) generation underground. Both diploid and haploid generations obligately mycorrhizal. The family Psilotaceae (whisk ferns, 2 genera), sister to Ophioglossaceae, is represented in California (SCo) by 1 (of 2 total) apparently introduced species, Psilotum nudum (L.) P. Beauv. Psilotum is easily distinguished by the dichotomously branching, almost leafless green stems, lack of roots, and large (2--3 mm) 3-lobed sporangia; sporangia are borne on the adaxial (upper) side of a minute (+- 1 mm) forked leaf. Pantrop, subtrop (nearest native populations in Arizona and in Sonora, Mexico); expected in cultivation areas, especially at bases of old palms, possibly brought in on root masses as subterranean gametophytes. [Pryer et al. 2004 Amer J Bot 91:1582--1598]
eFlora Treatment Author: Donald R. Farrar, except as noted
Scientific Editor: Alan R. Smith, Bruce G. Baldwin, Thomas J. Rosatti.
Genus: BotrychiumView DescriptionDichotomous Key

Common Name: MOONWORT
Habit: Roots smooth, pale yellow, without bulblets or plantlets. Leaf: deciduous; bud glabrous; sporophore and trophophore (or 2 sporophores) joined at or well above ground level; trophophore generally 1--2-pinnate (simple or entire or 0), linear to deltate to ternately triangular, thin to fleshy, pinnae ovate to oblong and midribbed or wedge- to fan-shaped and not midribbed, veins free, forked; sporophore 1--2-pinnate, rarely absent. Sporangia: not sunken in axis; stalk 0 or short.
Species In Genus: 25--35 species: generally temperate to arctic or alpine. Etymology: (Greek: bunch of grapes, from clusters of sporangia) Note: Difficult, needs study; most species uncommon, sporadic; good sampling of populations highly desirable in specimens, which must be carefully spread and pressed for identification. Botrychium multifidum moved to Sceptridium. Botrychium pedunculosum W.H. Wagner, differing from Botrychium pinnatum in having trophophore stalk +- = trophophore rachis (vs trophophore stalk 0 to 1/10 trophophore rachis), recently confirmed for California, based on discovery in summer of 2010 near Reynolds Creek, western of Yosemite National Park, Calaveras Co.

Botrychium pinnatum H. St. John
NATIVE
Leaf: sporophore, trophophore joined distal to mid-leaf; trophophore stalk 0--0.2 mm, blade pinnately lobed to 2-pinnate, 3--6 cm, 2--4 cm wide, oblong to deltate, pinnae 4--7 pairs, ovate to elliptic, shiny, bright green, midribbed, deeply lobed at base to entire near tip, margins entire to minutely crenate; sporophore 2-pinnate, deltate, stalk +- = trophophore. Chromosomes: 2n=180.
Ecology: Moist fields, shrubby slopes; Elevation: 1900--2800 m. Bioregional Distribution: KR (Etna Mills, Siskiyou Co.), CaRH (Mount Shasta, Domingo Lake se of Lassen Peak), c SNH (Bond Pass, Tuolumne Co.); Distribution Outside California: to Alaska, central Canada, Colorado. Note: A similar 2 ×-dissected moonwort, Botrychium lanceolatum (S.G. Gmel.) Angstr., occurs at Crater Lake, Oregon and may be expected in California. It differs from Botrychium pinnatum in having ternately dissected trophophore with narrow, pointed lobes.
Synonyms: Botrychium boreale J. Milde subsp. obtusilobum (Rupr.) R.T. Clausen
eFlora Treatment Author: Donald R. Farrar
Jepson Online Interchange
Listed on CNPS Rare Plant Inventory

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Citation for this treatment: Donald R. Farrar 2016. Botrychium pinnatum, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=15941, accessed on December 10, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on December 10, 2016.


Botrychium pinnatum
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© 1997 John Game
Botrychium pinnatum
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© 2007 John Game

More photos of Botrychium pinnatum in CalPhotos


Geographic subdivisions for Botrychium pinnatum:
KR (Etna Mills, Siskiyou Co.), CaRH (Mount Shasta, Domingo Lake se of Lassen Peak), c SNH (Bond Pass, Tuolumne Co.);
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.