Jepson eFlora: Taxon page
Key to families | Table of families and genera
Previous taxon Index to accepted names and synonyms:
| A | B | C | D | E | F | G | H | I | J | K | L | M | N | O | P | Q | R | S | T | U | V | W | X | Y | Z |
Next taxon
Berula erecta

Higher Taxonomy
Family: Apiaceae (Umbelliferae)View DescriptionDichotomous Key

Habit: Annual to perennial herb [shrub, tree], generally from taproot. Stem: generally +- scapose, generally ribbed, hollow. Leaf: basal and generally cauline, generally alternate; stipules generally 0; petiole base generally sheathing stem; blade generally much dissected, occasionally compound. Inflorescence: umbel or head, simple or compound, generally peduncled; bracts present in involucres or 0; bractlets generally present in "involucels". Flower: many, small, generally bisexual (or some staminate), generally radial (or outer bilateral); calyx 0 or lobes 5, small; petals 5, free, generally ovate or spoon-shaped, generally incurved at tips, generally +- ephemeral; stamens 5; pistil 1, ovary inferior, 2-chambered, generally with a +- conic, persistent projection or platform at tip subtending 2 free styles. Fruit: 2 dry, 1-seeded halves (= mericarps), separating from each other but generally +- persistent to central axis; ribs on halves 5, 2 marginal, 3 to back; oil tubes 1--several per interval between ribs.
Genera In Family: 300 genera, 3000 species: +- worldwide, especially temperate; many cultivated for food or spice (e.g., Carum, caraway; Daucus; Petroselinum); Bupleurum lancifolium Hornem. is historical garden weed; some toxic (e.g., Conium). Note: Mature fruit generally critical in identification, shape given in outline. Hydrocotyle moved to Araliaceae. Petroselinum crispum (Mill.) A.W. Hill is a waif.
eFlora Treatment Author: Lincoln Constance & Margriet Wetherwax, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.
Genus: BerulaView Description 


Species In Genus: 1 sp. Etymology: (Latin: water-cress)

Berula erecta (Huds.) Coville
Habit: Perennial herb, < 15 dm, stoloned, glabrous. Stem: ascending or erect, branched, hollow, rooting at proximal nodes. Leaf: petiole 4--12 cm, narrowly sheathing; blade 1.5 dm, oblong, 1-pinnate, much dissected if submerged, leaflets 7--12 pairs, 1--8 cm, oblong to ovate, sessile, lobed or serrate. Inflorescence: umbels compound, peduncled, terminal or opposite leaves; bracts, bractlets leaf-like, lanceolate; rays, pedicels +- unequal, spreading. Flower: calyx lobes minute, persistent; petals wide, white. Fruit: 1.5--2 mm, +- round, compressed side-to-side, glabrous; ribs thread-like, inconspicuous in corky fruit wall; oil tubes apparent, deeply embedded in fruit wall; fruit axis divided to base, branches adhering to and falling with fruit-halves. Chromosomes: 2n=18.
Ecology: Marshy areas, streams; Elevation: < 2000 m. Bioregional Distribution: CA; Distribution Outside California: to British Columbia, New York, Texas, Mexico; Eurasia, Africa. Toxicity: Possibly TOXIC: possibly poisonous to livestock. Flowering Time: Jul--Oct
eFlora Treatment Author: Lincoln Constance & Margriet Wetherwax
Jepson Online Interchange

Previous taxon: Berula
Next taxon: Bowlesia

Name Search
botanical illustration including Berula erecta


Citation for this treatment: Lincoln Constance & Margriet Wetherwax 2016. Berula erecta, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 05, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on May 05, 2016.

Berula erecta
click for enlargement
© 2014 Neal Kramer
Berula erecta
click for enlargement
© 2006 Keir Morse
Berula erecta
click for enlargement
© 2014 Neal Kramer
Berula erecta
click for enlargement
© 2012 Steve Matson
Berula erecta
click for enlargement
© 2014 Neal Kramer
Berula erecta
click for enlargement
© 2015 Steve Matson

More photos of Berula erecta in CalPhotos

Geographic subdivisions for Berula erecta:
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

View elevation by latitude chart
Data provided by the participants of the Consortium of California Herbaria.
View all CCH records

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.