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Astragalus layneae
LAYNE MILKVETCH

Higher Taxonomy
Family: Fabaceae (Leguminosae)View DescriptionDichotomous Key

Common Name: LEGUME FAMILY
Habit: Annual to tree. Leaf: generally alternate, generally compound, generally stipuled, generally entire, pinnately veined Inflorescence: generally raceme, spike, umbel or head; or flowers 1--few in axils. Flower: generally bisexual, generally bilateral; hypanthium 0 or flat to tubular; sepals generally 5, generally fused; petals generally 5, free, fused, or lower 2 +- united into keel (see 3, Key to Groups, for banner, wings); stamens 10 or many (or [1], 5, 6, 7, 9), free or fused or 10 with 9 filaments at least partly fused, 1 (uppermost) free; pistil 1, ovary superior, generally 1-chambered, ovules 1--many, style, stigma 1. Fruit: legume, including a stalk-like base (above receptacle) or not. Seed: 1--many, often +- reniform, generally hard, smooth.
Genera In Family: +- 730 genera, 19400 species: worldwide; with grasses, requisite in agriculture, most natural ecosystems. Many cultivated, most importantly Arachis, peanut; Glycine, soybean; Phaseolus, beans; Medicago, alfalfa; Trifolium, clovers; many orns. Note: Unless stated otherwise, fruit length including stalk-like base, number of 2° leaflets is per 1° leaflet. Upper suture of fruit adaxial, lower abaxial. Anthyllis vulneraria L. evidently a waif, a contaminant of legume seed from Europe. Laburnum anagyroides Medik., collected on Mount St. Helena in 1987, may be naturalized. Ceratonia siliqua L., carob tree (Group 2), differs from Gleditsia triacanthos L. in having evergreen (vs deciduous) leaves that are 1-pinnate (vs 1-pinnate on spurs on old stems, 2-pinnate on new stems) with 2--5(8) (vs 7--17) 1° leaflets, commonly cultivated, now naturalized in southern California. Aeschynomene rudis Benth. , Halimodendron halodendron (Pall.) Voss (possibly extirpated), Lens culinaris Medik. are agricultural weeds. Caragana arborescens Lam. only cult. Ononis alopecuroides L. , Sphaerophysa salsula (Pall.) DC. all evidently extirpated. Cercidium moved to Parkinsonia; Chamaecytisus to Cytisus; Psoralidium lanceolatum to Ladeania.
eFlora Treatment Author: Martin F. Wojciechowski, except as noted
Scientific Editor: Martin F. Wojciechowski, Thomas J. Rosatti.
Genus: AstragalusView DescriptionDichotomous Key

Common Name: LOCOWEED, MILKVETCH
Habit: Annual, perennial herb from crown, generally unarmed; hairs generally present, simple or branches 2, from base, parallel to leaf surface, unequal or not. Stem: 0 or prostrate to erect. Leaf: odd-1-pinnate (or palmately compound); leaflets generally jointed to midrib, entire; stipules membranous, lower fused around stem into sheaths (stipule sheaths) or not. Inflorescence: raceme, head- or umbel-like or not, axillary; flowers 2--many. Flower: bilateral; keel petals with small protrusion at base locking into pit on adjacent wing; 9 filaments fused, 1 free; ovary (and fruit) generally sessile, style slender, stigma minute. Fruit: generally 1- or +- 2-chambered, often mottled, generally +- dry in age, sometimes deciduous (falling from plant with or without pedicel, calyx, receptacle) before dehiscence. Seed: 2--many, smooth, compressed, +- notched at attachment scar.
Species In Genus: > 2500 species: +- worldwide (380 in North America, 97 in California, including many rare taxa). Etymology: (Greek: ankle-bone or dice, perhaps from rattling of seeds within fruit) Note: Difficult; flower and fruit needed for identification; fruit said to be "deciduous" dehisce only after fruit has separated from pl; many good species appear similar; some species complexes need study. Taxa near province boundaries may appear in > 1 key. Varieties keyed under species for simplicity; species with varieties so identified in key. Fruit length including beak and any stalk-like base unless fruit body specified; fruit depth is suture-to-suture axis. Astragalus tephrodes A. Gray var. brachylobus (A. Gray) Barneby in southwestern Utah, Arizona, near California.
eFlora Treatment Author: Martin F. Wojciechowski & Richard Spellenberg

Astragalus layneae Greene
NATIVE
Habit: Perennial herb from deep rhizome (generally overlooked); hairs coarse, generally +- gray. Stem: erect, 2.5--16 cm. Leaf: 4--16 cm; leaflets 11--23, 5--23 mm, ovate to +- round. Inflorescence: flowers 10--45, early ascending, then spreading. Flower: calyx hairs mostly black, some white; petals +- white with keel tip, wing tips purple, banner often lilac-tinged, banner 12.5--18 mm, recurved +- 50°, keel 10.4--16.5 mm. Fruit: 20--65 mm, 3.5--8 mm wide, incurved 1/4--full circle, leathery, hairs spreading, wavy; chambers +- 2 in lower 1/2. Chromosomes: 2n=44.
Ecology: Sandy flats, washes; Elevation: 25--1750 m. Bioregional Distribution: W&I, DMoj; Distribution Outside California: southern Nevada, northwestern Arizona. Flowering Time: Mar--Jun
eFlora Treatment Author: Martin F. Wojciechowski & Richard Spellenberg
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Citation for this treatment: Martin F. Wojciechowski & Richard Spellenberg 2016. Astragalus layneae, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=14886, accessed on September 28, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on September 28, 2016.


Astragalus layneae
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© 2008 Neal Kramer
Astragalus layneae
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© 2006 Gary A. Monroe
Astragalus layneae
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© 2014 Neal Kramer
Astragalus layneae
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© 2009 Gary A. Monroe
Astragalus layneae
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© 2010 Aaron Schusteff
Astragalus layneae
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© 2010 Aaron Schusteff

More photos of Astragalus layneae in CalPhotos


Geographic subdivisions for Astragalus layneae:
W&I, DMoj;
Markers link to CCH specimen records. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues. Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.