Jepson eFlora: Taxon page
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Anulocaulis annulatus

Higher Taxonomy
Family: NyctaginaceaeView DescriptionDichotomous Key

Habit: Annual, perennial herb, subshrub, [shrub, tree], glabrous or hairy. Stem: often forked. Leaf: opposite, sessile or petioled, pairs generally unequal; blade generally entire. Inflorescence: generally forked; spike, head-like cluster, or umbel, flowers rarely 1, bracts forming a calyx-like involucre or not. Flower: bisexual, generally +- radial (bilateral), sometimes cleistogamous in some genera; perianth of 1 whorl, generally petal-like, bell- to trumpet-shaped, base hardened, tightly surrounding ovary in fruit, lobes 3--5, generally notched to +- lobed; stamens 1--many; ovary superior (appearing inferior due to hardened perianth base), style 1. Fruit: achene in hardened perianth base; round to +- flat; smooth, angled, ribbed, or winged; glabrous, hairy, or glandular.
Genera In Family: 30 genera, 350 species: warm regions, especially America; some ornamental (Bougainvillea; Mirabilis, four o'clock).
eFlora Treatment Author: Andy Murdock, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: AnulocaulisView Description 

Common Name: RINGSTEM
Habit: Perennial herb from thick caudex. Stem: little-branched, erect, > 7 mm diam; internodes with sticky brown ring. Leaf: few, +- in lower 1/2, petioled; blade oblong to round, thick. Inflorescence: openly branched; flowers in head, raceme, or umbel-like cluster; bracts 1--3, free, not forming involucre. Flower: perianth funnel-shaped; stamens 3, exserted; stigma +- spheric, exserted. Fruit: inconspicuously 10-ribbed, glabrous.
Species In Genus: 5 species: especially Chihuahuan Desert, northeastern Mexico. Etymology: (Latin: ring stem, from sticky ring on internode) Note: Closely related to Boerhavia.
eFlora Treatment Author: Andy Murdock & Richard Spellenberg

Anulocaulis annulatus (Coville) Standl.
Stem: < 1.5 m. Leaf: blade 3--10 cm, oblong to round-reniform, hairs stiff, with enlarged, dark, glandular base. Inflorescence: terminal, panicle-like, 1--2 branches per node, each ending in head-like cluster of flowers; bracts leaf-like, reduced; peduncle 15--40 mm. Flower: perianth +- 8 mm, tube +- green, hairy, limb pale pink. Fruit: 4--5 mm, spindle-shaped, gray-brown. Chromosomes: 2n=48.
Ecology: Rocky slopes, canyons; Elevation: < 1200 m. Bioregional Distribution: ne DMoj (Death Valley region). Flowering Time: Apr--May
Synonyms: Boerhavia annulatus Coville
eFlora Treatment Author: Andy Murdock & Richard Spellenberg
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botanical illustration including Anulocaulis annulatus


Citation for this treatment: Andy Murdock & Richard Spellenberg 2016. Anulocaulis annulatus, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 01, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on May 01, 2016.

Anulocaulis annulatus
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© 2016 Steve Matson
Anulocaulis annulatus
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© 2010 Barry Breckling
Anulocaulis annulatus
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© 2016 Steve Matson
Anulocaulis annulatus
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© 2009 Gary A. Monroe
Anulocaulis annulatus
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© 2005 Steve Matson
Anulocaulis annulatus
click for enlargement
© 2009 Gary A. Monroe

More photos of Anulocaulis annulatus in CalPhotos

Geographic subdivisions for Anulocaulis annulatus:
ne DMoj (Death Valley region).
Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

View elevation by latitude chart
Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.