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Galium
BEDSTRAW

Higher Taxonomy
Family: RubiaceaeView DescriptionDichotomous Key

Common Name: MADDER FAMILY
Habit: Annual to tree, vine. Leaf: generally opposite (whorled), entire; stipules generally fused to stem, adjacent pairs occasionally fused, or occasionally leaf-like and appearing like whorled leaves. Inflorescence: cyme, panicle, spike, cluster, or flower 1, generally terminal and +- axillary. Flower: generally bisexual; calyx +- 4(5)-lobed, occasionally 0 (Galium, Crucianella) or 6 (Sherardia); corolla generally radial, 4(5)-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1(2). Fruit: drupe, berry, or 2 or 4 nutlets [capsule].
Genera In Family: +- 500 genera, 6000 species: worldwide, especially tropics; many cultivated, including Coffea, coffee; Cinchona, quinine; many ornamental. Note: Diodia teres Walter doubtfully in California.
eFlora Treatment Author: Robert E. Preston & Lauramay T. Dempster, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.

Galium
Habit: Annual, perennial herb, occasionally subshrub, glabrous to hairy, generally scabrous; dioecious, bisexual, or flowers unisexual and bisexual. Stem: 4-angled, occasionally ridged lengthwise. Leaf: generally in whorls of >= 4, including leaf-like stipules. Inflorescence: panicles, axillary clusters (cymes), or occasionally 1 in axils. Flower: bisexual, or unisexual with sterile stamens or pistils; calyx 0; corolla generally rotate, occasionally +- bell-shaped, generally +- green, yellow to white, occasionally pink or red, lobes generally 4; ovary 2-lobed, styles 2, bases +- fused. Fruit: 2 nutlets or berry.
Species In Genus: +- 650 species: worldwide, especially temperate. Etymology: (Greek: milk, from use of some species for curdling) Note: Ovary and fruit generally +- equally hairy on a pl; staminate plants generally identified by vestigial ovaries, pistillate plants generally by vestigial anthers. Galium saxatile L., Galium schultesii Vest, and Galium verum L. are lawn weeds in California.
eFlora Treatment Author: Valerie Soza
Jepson Online Interchange
Key to Galium

Previous taxon: Diodia teres
Next taxon: Galium ambiguum

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Citation for this treatment: Valerie Soza 2016. Galium, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=11284, accessed on August 27, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on August 27, 2016.


Galium angustifolium subsp. foliosum
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© 2014 Steve Matson
Galium angustifolium subsp. nudicaule
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© 2015 Keir Morse
Galium bolanderi
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© 2009 Barry Breckling
Galium catalinense subsp. acrispum
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© 2011 Neal Kramer
Galium johnstonii
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© 2015 Keir Morse
Galium obtusum subsp. obtusum
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© 2016 Keir Morse

More photos of Galium in CalPhotos