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Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key

Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera; treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.

Habit: Perennial herb (biennial); hairs appressed to spreading; caudex generally branched in age, often +- woody, taprooted. Stem: ascending or erect. Leaf: lowest petioles generally +- = blades, +- winged, others 0. Inflorescence: coiled cymes, generally > 3, generally terminal and axillary, +- bracted; pedicel in fruit elongated, recurved to reflexed. Flower: calyx deep-5-lobed; corolla rotate-salverform, generally white with yellow patch adaxially, lobes appendaged near base. Fruit: nutlets erect, > style, attachment scar lateral-medial, generally with barb-tipped prickles abaxially and on margin.
Species In Genus: 40 species: generally western North America, southeastern Asia. Etymology: (J. Hackel, Czech botanist, 1783--1869) Note: Values for corolla limb diam take into account shrinkage during flower period. Difficult, study needed, especially in northern California, southeastern Asia; sometimes merged with Lappula.
Unabridged Note: In North America filling old-world role of flat-flowered forget-me-nots, Myosotis, Lepechiniella, etc, for pollinators.
eFlora Treatment Author: Ronald B. Kelley & Robert L. Carr
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Citation for this treatment: Ronald B. Kelley & Robert L. Carr 2016. Hackelia, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on October 20, 2016.

Citation for the whole project: Jepson Flora Project (eds.) 2016. Jepson eFlora,, accessed on October 20, 2016.

Hackelia brevicula
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© 2015 Steve Matson
Hackelia nervosa
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© 2003 Christopher L. Christie
Hackelia micrantha
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© 2010 Keir Morse
Hackelia velutina
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© 2015 Barry Rice
Hackelia micrantha
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© 2009 Keir Morse
Hackelia micrantha
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© 2004 Steve Matson

More photos of Hackelia in CalPhotos