trait.id trait state.id dstate
1 Growth form 1 short basal
1 Growth form 2 long basal
1 Growth form 3 semi-basal
1 Growth form 4 erect leafy
1 Growth form 5 cushions (=pulvinate)
1 Growth form 6 tussocks
1 Growth form 7 dwarf shrubs
1 Growth form 8 shrubs
1 Growth form 9 trees
1 Growth form 10 leafless shrubs or trees
1 Growth form 11 short succulents
1 Growth form 12 tall succulents
1 Growth form 13 palmoids
1 Growth form 14 epiphytes
1 Growth form 15 climbers and scramblers
1 Growth form 16 hemi-epiphytes
1 Growth form 17 hemiparasites or holoparasites
1 Growth form 18 aquatic submerged
1 Growth form 19 aquatic floating
1 Growth form 20 other growth forms.
2 Life form 1 phanerophytes
2 Life form 2 chamaephytes
2 Life form 3 hemicryptophytes
2 Life form 4 geophytes
2 Life form 5 therophytes
2 Life form 6 helophytes
2 Life form 7 hydrophytes
4 Plant habit 1 winter annual
4 Plant habit 2 spring annual
4 Plant habit 3 summer annual
4 Plant habit 4 perennial herbaceous
4 Plant habit 5 shrub
4 Plant habit 6 tree
5 Maximum height (d) 1 0-3 cm
5 Maximum height (d) 2 3-10 cm
5 Maximum height (d) 3 10-50
5 Maximum height (d) 4 50-100
5 Maximum height (d) 5 100-150
5 Maximum height (d) 6 150-200
5 Maximum height (d) 7 200-300
5 Maximum height (d) 8 >300
7 Flower height (d) 1 0-5 cm
7 Flower height (d) 2 5-10
7 Flower height (d) 3 10-50
7 Flower height (d) 4 50-100
7 Flower height (d) 5 100-150
7 Flower height (d) 6 150-200
7 Flower height (d) 7 200-300
7 Flower height (d) 8 >300
9 Dead branch retention 1 no
9 Dead branch retention 2 some
9 Dead branch retention 3 a lot
57 Twig dry matter content 1 <200 mg g-1
57 Twig dry matter content 2 200–400
57 Twig dry matter content 3 400–600
57 Twig dry matter content 4 600–800
57 Twig dry matter content 5 >800
58 Twig drying time 1 ²5 days
58 Twig drying time 2 4
58 Twig drying time 3 3
58 Twig drying time 4 2
58 Twig drying time 5 ²1
10 Degree of ramification (branching) 1 no branches
10 Degree of ramification (branching) 2 only 1st order
10 Degree of ramification (branching) 3 2 orders
10 Degree of ramification (branching) 4 3 orders
10 Degree of ramification (branching) 5 4 orders
11 Standing fine litter in driest season 1 none
11 Standing fine litter in driest season 2 some
11 Standing fine litter in driest season 3 substantial (with dead leaves/twigs or flaking bark)
11 Standing fine litter in driest season 4 more dead than live fine mass aboveground
11 Standing fine litter in driest season 5 shoot dies back entirely
11 Standing fine litter in driest season 6 standing as one litter unit
12 Volatile oils, waxes and/or resins 1 none
12 Volatile oils, waxes and/or resins 2 some
12 Volatile oils, waxes and/or resins 3 substantial
12 Volatile oils, waxes and/or resins 4 abundant
12 Volatile oils, waxes and/or resins 5 very abundant
13 Leaf type 1 simple
13 Leaf type 2 compound
13 Leaf type 3 dissected or lobed
13 Leaf type 4 modified
13 Leaf type 5 absent
14 Compound leaf type 1 palmate
14 Compound leaf type 2 pinnate
14 Compound leaf type 3 bipinnate
14 Compound leaf type 4 odd
14 Compound leaf type 5 even
15 Blade morphology 1 cylindric
15 Blade morphology 2 scale-like
15 Blade morphology 3 narrowly linear
15 Blade morphology 4 linear
15 Blade morphology 5 ovate
15 Blade morphology 6 elliptic
15 Blade morphology 7 deltate
15 Blade morphology 8 cordate
15 Blade morphology 9 reniform
16 Blade length (d) 1 <0.01 cm
16 Blade length (d) 2 0.01-1
16 Blade length (d) 3 1-5
16 Blade length (d) 4 5-10
16 Blade length (d) 5 10-50
16 Blade length (d) 6 50-100
16 Blade length (d) 7 >100
17 Blade width (d) 1 <0.01 cm
17 Blade width (d) 2 0.01-1
17 Blade width (d) 3 1-5
17 Blade width (d) 4 5-10
17 Blade width (d) 5 10-50
17 Blade width (d) 6 >50
20 Compound leaf length (d) 1 <0.01 cm
20 Compound leaf length (d) 2 0.01-1
20 Compound leaf length (d) 3 1-5
20 Compound leaf length (d) 4 5-10
20 Compound leaf length (d) 5 10-50
20 Compound leaf length (d) 6 50-100
20 Compound leaf length (d) 7 100-500
20 Compound leaf length (d) 7 >500
21 Compound leaf width (d) 1 <0.01 cm
21 Compound leaf width (d) 2 0.01-1
21 Compound leaf width (d) 3 1-5
21 Compound leaf width (d) 4 5-10
21 Compound leaf width (d) 5 10-50
21 Compound leaf width (d) 6 50-100
21 Compound leaf width (d) 7 >100
25 Blade area (d) 1 <0.01 cm2
25 Blade area (d) 2 0.01-1
25 Blade area (d) 3 1-5
25 Blade area (d) 4 5-10
25 Blade area (d) 5 10-50
25 Blade area (d) 6 50-100
25 Blade area (d) 7 > 100
27 Raunkiaer leaf size 1 leptophyll < 25 mm2
27 Raunkiaer leaf size 2 nanophyll 25-225
27 Raunkiaer leaf size 3 microphyll 225-2025
27 Raunkiaer leaf size 4 mesophyll 2025-18225
27 Raunkiaer leaf size 5 macropyll 18225-164025
27 Raunkiaer leaf size 6 megaphyll > 164026.
28 Blade margin 1 not applicable
28 Blade margin 2 entire
28 Blade margin 3 crenate
28 Blade margin 4 dentate
28 Blade margin 5 lobed
28 Blade margin 6 lobed and dentate
28 Blade margin 7 serrate
33 Stomatal distribution 1 both
33 Stomatal distribution 2 lower
33 Stomatal distribution 3 upper
33 Stomatal distribution 4 none
33 Stomatal distribution 5 stem
39 Leaf texture 1 sclerophyll
39 Leaf texture 2 mesophyll
39 Leaf texture 3 succulent
54 Leaf habit 1 evergreen
54 Leaf habit 2 summer deciduous
54 Leaf habit 3 winter deciduous
54 Leaf habit 4 semi-deciduous
55 Photosynthetic pathway 1 C3
55 Photosynthetic pathway 2 C4
55 Photosynthetic pathway 3 CAM
59 Green/photosynthetic stems 1 yes
59 Green/photosynthetic stems 2 no
60 Stem succulence 1 yes
60 Stem succulence 2 no
60 Stem succulence 3 in other organs (please specify)
61 Bark thickness 1 less 1mm
61 Bark thickness 2 1-10
61 Bark thickness 3 >10
63 Bark surface structure (texture) 1 smooth
63 Bark surface structure (texture) 2 very slight texture
63 Bark surface structure (texture) 3 intermediate texture
63 Bark surface structure (texture) 4 strong texture
63 Bark surface structure (texture) 5 very coarse texture.
64 Spinescence (density) 1 no spines, thorns or prickles
64 Spinescence (density) 2 low or very local density of soft spine equivalents <5 mm;
64 Spinescence (density) 3 high density of soft spine equivalents or intermediate density of spine equivalents of intermediate hardness;
64 Spinescence (density) 4 intermediate or high density of hard sharp spine equivalents >5 mm;
64 Spinescence (density) 5 intermediate or high density of hard sharp spine equivalents >20 mm;
64 Spinescence (density) 6 intermediate or high density of hard sharp spine equivalents >100 mm
65 Surface indumenta on leaves 1 glabrous
65 Surface indumenta on leaves 2 canescent
65 Surface indumenta on leaves 3 ciliate
65 Surface indumenta on leaves 4 prickle
65 Surface indumenta on leaves 5 puberulent
65 Surface indumenta on leaves 6 scabrous
65 Surface indumenta on leaves 7 scale
65 Surface indumenta on leaves 8 strigose
65 Surface indumenta on leaves 9 tomentose.
66 Surface indumenta on stems 1 glabrous
66 Surface indumenta on stems 2 canescent
66 Surface indumenta on stems 3 ciliate
66 Surface indumenta on stems 4 prickle
66 Surface indumenta on stems 5 puberulent
66 Surface indumenta on stems 6 scabrous
66 Surface indumenta on stems 7 scale
66 Surface indumenta on stems 8 strigose
66 Surface indumenta on stems 9 tomentose.
67 Surface indumenta on flowers 1 glabrous
67 Surface indumenta on flowers 2 canescent
67 Surface indumenta on flowers 3 ciliate
67 Surface indumenta on flowers 4 prickle
67 Surface indumenta on flowers 5 puberulent
67 Surface indumenta on flowers 6 scabrous
67 Surface indumenta on flowers 7 scale
67 Surface indumenta on flowers 8 strigose
67 Surface indumenta on flowers 9 tomentose.
68 Surface indumenta on fruits 1 glabrous
68 Surface indumenta on fruits 2 canescent
68 Surface indumenta on fruits 3 ciliate
68 Surface indumenta on fruits 4 prickle
68 Surface indumenta on fruits 5 puberulent
68 Surface indumenta on fruits 6 scabrous
68 Surface indumenta on fruits 7 scale
68 Surface indumenta on fruits 8 strigose
68 Surface indumenta on fruits 9 tomentose.
69 Surface indumenta on buds 1 glabrous
69 Surface indumenta on buds 2 canescent
69 Surface indumenta on buds 3 ciliate
69 Surface indumenta on buds 4 prickle
69 Surface indumenta on buds 5 puberulent
69 Surface indumenta on buds 6 scabrous
69 Surface indumenta on buds 7 scale
69 Surface indumenta on buds 8 strigose
69 Surface indumenta on buds 9 tomentose.
70 Aromatic/Chemical defences 1 not present or present in not significant amounts,
70 Aromatic/Chemical defences 2 terpenoids (pyrethrum, essential oils, resins, latex)
70 Aromatic/Chemical defences 3 phenolics (lignins, tannins, flavonoids, allelopathics)
70 Aromatic/Chemical defences 4 alkaloids
70 Aromatic/Chemical defences 5 present, type unknown
71 Root depth: 1. type 1 shallow
71 Root depth: 1. type 2 deep
72 Root depth: 2. depth 1 <0.25 m
72 Root depth: 2. depth 2 0.25-0.5
72 Root depth: 2. depth 3 0.5-1
72 Root depth: 2. depth 4 1-2
72 Root depth: 2. depth 5 2-5
72 Root depth: 2. depth 6 5-10
72 Root depth: 2. depth 7 >10
74 Root type 1 absent
74 Root type 2 tap root
74 Root type 3 fibrous
74 Root type 4 adventitious
74 Root type 5 contractile
75 Mycorrhizal association 1 arbuscular mycorrhiza
75 Mycorrhizal association 2 ectomycorrhiza
75 Mycorrhizal association 3 ericoid
75 Mycorrhizal association 4 orchid
75 Mycorrhizal association 5 no mycorrhizal association
75 Mycorrhizal association 6 unknown
76 Strength of mycorrhizal association 1 obligate or nearly obligate
76 Strength of mycorrhizal association 2 facultative
76 Strength of mycorrhizal association 3 not important
76 Strength of mycorrhizal association 4 unknown
77 N fixation 1 Rhizobium
77 N fixation 2 Actinorhiza (Frankia)
77 N fixation 3 No
79 Seed longevity 1 transient (<1 year)
79 Seed longevity 2 short-term persistent (1-5 years)
79 Seed longevity 3 long-term persistent (>5 years)
80 Seed length 1 <1 mm
80 Seed length 2 1-5
80 Seed length 3 5-10
80 Seed length 4 10-25
80 Seed length 5 25-50
80 Seed length 6 50-100
80 Seed length 7 >100
81 Seed width 1 <1 mm
81 Seed width 2 1-5
81 Seed width 3 5-10
81 Seed width 4 10-25
81 Seed width 5 25-50
81 Seed width 6 50-100
81 Seed width 7 >100
83 Annual seed production per plant 1 1-10 seeds/plant
83 Annual seed production per plant 2 10-100
83 Annual seed production per plant 3 100-1000
83 Annual seed production per plant 4 1000-10000
83 Annual seed production per plant 5 >10000
85 Heterocarpy 1 yes
85 Heterocarpy 2 no
86 Dispersal mode 1 unassisted dispersal
86 Dispersal mode 2 wind dispersal (anemochory)
86 Dispersal mode 3 internal animal transport (endo-zoochory)
86 Dispersal mode 4 external animal transport (exo-zoochory)
86 Dispersal mode 5 dispersal by hoarding
86 Dispersal mode 6 ant dispersal (myrmecochory)
86 Dispersal mode 7 dispersal by water (hydrochory)
86 Dispersal mode 8 dispersal by launching (ballistichory)
86 Dispersal mode 9 bristle contraction
87 Diaspore type 1 vegetative dispersule
87 Diaspore type 2 one-seeded generative dispersule
87 Diaspore type 3 multi-seeded generative dispersule
88 Seed structure 1 Nutrient rich appendages or structures: Elaisome
88 Seed structure 2 Nutrient rich appendages or structures: Aril
88 Seed structure 3 Nutrient rich appendages or structures: Pulp
88 Seed structure 4 Balloon structures: Open structures
88 Seed structure 5 Balloon structures: Closed structures
88 Seed structure 6 Small flat appendages
88 Seed structure 7 Large flat appendages
88 Seed structure 8 One short elongated appendage
88 Seed structure 9 Two or more short appendages
88 Seed structure 10 One long elongated appendage
88 Seed structure 11 Two or more long elongated appendages
88 Seed structure 12 Hooked
88 Seed structure 13 Other specializations
88 Seed structure 14 No appendages: Structured surface
88 Seed structure 15 No appendages: Smooth surface
88 Seed structure 7 Unknown
89 Fruit morphology 1 achene
89 Fruit morphology 2 berry
89 Fruit morphology 3 capsule
89 Fruit morphology 4 drupe
89 Fruit morphology 5 follicle
89 Fruit morphology 6 legume
89 Fruit morphology 7 nut
89 Fruit morphology 8 pome
89 Fruit morphology 9 utricle
90 Dehiscent fruit 1 yes
90 Dehiscent fruit 2 no
91 Simple fruit 1 yes
91 Simple fruit 2 no (compound)
92 Fleshy fruit 1 yes
92 Fleshy fruit 2 no
93 Number of seeds per fruit 1 1 seed
93 Number of seeds per fruit 2 1-3
93 Number of seeds per fruit 3 3-5
93 Number of seeds per fruit 4 5-10
93 Number of seeds per fruit 5 10-15
93 Number of seeds per fruit 6 15-30
93 Number of seeds per fruit 7 30-50
93 Number of seeds per fruit 8 50-100
93 Number of seeds per fruit 9 100-500
93 Number of seeds per fruit 10 500-1000
93 Number of seeds per fruit 11 >1000
94 Germination (seed type) 1 Recalcitrant
94 Germination (seed type) 2 Orthodox
94 Germination (seed type) 3 Dormant
94 Germination (seed type) 4 Non-dormant
95 Germination requirements 1 high temperature
95 Germination requirements 2 low temperature
95 Germination requirements 3 fluctuating temperature
95 Germination requirements 4 drying at high temperature
95 Germination requirements 5 darkness
95 Germination requirements 6 light
95 Germination requirements 7 gases: oxygen, carbon dioxide, ethylene, etc..
95 Germination requirements 8 water: hydratation, wet-dry cycles, etc..
95 Germination requirements 9 inorganic chemicals: nitrate nitrite, etc.
95 Germination requirements 10 fire
95 Germination requirements 11 smoke
95 Germination requirements 12 charate
95 Germination requirements 13 animal digestive system
95 Germination requirements 14 unknown
96 Longevity 1 <1 yr
96 Longevity 2 1-5
96 Longevity 3 6-100
96 Longevity 4 100+
97 Reproductive episodes 1 semelparous
97 Reproductive episodes 2 iteroparous
98 Minimum age at reproductive maturity 1 <1 yr
98 Minimum age at reproductive maturity 2 1-2
98 Minimum age at reproductive maturity 3 3-5
98 Minimum age at reproductive maturity 4 6-10
98 Minimum age at reproductive maturity 5 10+
99 Clonality 1 non-clonal
99 Clonality 2 above-ground: stolons
99 Clonality 3 above-ground: gemmiparous
99 Clonality 4 above-ground: other vegetative buds or plant fragments
99 Clonality 5 below-ground: rhizomes
99 Clonality 6 below-ground: tubers
99 Clonality 7 below-ground: bulbs
99 Clonality 8 below-ground: adventitious root buds
100 Resprouting behavior 1 non-resprouting
100 Resprouting behavior 2 epicormic
100 Resprouting behavior 3 basal
100 Resprouting behavior 4 rhizome/root/lignotuber/burl
100 Resprouting behavior 5 unknown
101 Resprouting factors 1 herbivory
101 Resprouting factors 2 fire
101 Resprouting factors 3 wind
101 Resprouting factors 4 other factors (please specify)
102 Pollen vector 1 wind
102 Pollen vector 2 water
102 Pollen vector 3 pseudocleistogamy
102 Pollen vector 4 cleistogamy
102 Pollen vector 5 insects
102 Pollen vector 6 bats
102 Pollen vector 7 birds
102 Pollen vector 8 slugs
102 Pollen vector 9 other animals
102 Pollen vector 10 other abiotic vectors
103 Mating system 1 self-compatible
103 Mating system 2 ± self-compatible
103 Mating system 3 ± self-incompatible
103 Mating system 4 self-incompatible
104 Floral rewards 1 nectar
104 Floral rewards 2 pollen
104 Floral rewards 3 oil
104 Floral rewards 4 deceit
105 Flower size 1 < 5 mm
105 Flower size 2 5-10
105 Flower size 3 10-30
105 Flower size 4 30-50
105 Flower size 5 50-100
105 Flower size 6 100-200
105 Flower size 7 200-500
105 Flower size 8 > 500
108 Floral symmetry 1 asymmetric
108 Floral symmetry 2 bilateral
108 Floral symmetry 3 biradial
108 Floral symmetry 4 radial
109 Flower color 1 blue
109 Flower color 2 brown
109 Flower color 3 yellow
109 Flower color 4 green
109 Flower color 5 lilac
109 Flower color 6 orange
109 Flower color 7 purple
109 Flower color 8 red
109 Flower color 9 pink
109 Flower color 10 various colors
109 Flower color 11 violet
109 Flower color 12 white
110 Heterostyly 1 distyly
110 Heterostyly 2 tristily
110 Heterostyly 3 stigma-height dimorphism
110 Heterostyly 4 homostyly
111 Breeding system 1 allogamous
111 Breeding system 2 facultative allogamous
111 Breeding system 3 mixed mating
111 Breeding system 4 facultative autogamous
111 Breeding system 5 autogamous
111 Breeding system 6 automixis
112 Plant sex expression 1 hermaphrodite
112 Plant sex expression 2 monoecious
112 Plant sex expression 3 dioecious
112 Plant sex expression 4 gynomonoecious
112 Plant sex expression 5 andromonoecious
112 Plant sex expression 6 trimonoecious (polygamous).
120 Typical habitat 1 Coastal Strand
120 Typical habitat 2 Coastal Salt Marsh
120 Typical habitat 3 Freshwater Marsh
120 Typical habitat 4 Northern Coastal Scrub
120 Typical habitat 5 Coastal Sage Scrub
120 Typical habitat 6 Sagebrush Scrub
120 Typical habitat 7 Shadscale Scrub
120 Typical habitat 8 Creosote Bush Scrub
120 Typical habitat 9 Alkali Sink
120 Typical habitat 10 North Coastal Coniferous Forest
120 Typical habitat 11 Closed-cone Pine Forest
120 Typical habitat 12 Redwood Forest
120 Typical habitat 13 Douglas-Fir Forest
120 Typical habitat 14 Yellow Pine Forest
120 Typical habitat 15 Red Fir Forest
120 Typical habitat 16 Lodgepole Forest
120 Typical habitat 17 Subalpine Forest
120 Typical habitat 18 Bristle-cone Pine Forest
120 Typical habitat 19 Mixed Evergreen Forest
120 Typical habitat 20 Northern Oak Woodland
120 Typical habitat 21 Southern Oak Woodland
120 Typical habitat 22 Foothill Woodland
120 Typical habitat 23 Chaparral
120 Typical habitat 24 Coastal Prairie
120 Typical habitat 25 Valley Grassland
120 Typical habitat 26 Alpine Fell-field
120 Typical habitat 27 Northern Juniper Woodland
120 Typical habitat 28 Pinyon-Juniper Woodland
120 Typical habitat 29 Joshua Tree Woodland
125 Rabinowitz rarity: geographic 1 wide
125 Rabinowitz rarity: geographic 2 narrow
126 Rabinowitz rarity: habitat specificity 1 broad
126 Rabinowitz rarity: habitat specificity 2 restricted
127 Rabinowitz rarity: local abundance 1 somewhere large
127 Rabinowitz rarity: local abundance 2 everywhere small
128 Fed rarity status 1 Threatened
128 Fed rarity status 2 Endangered
128 Fed rarity status 3 Without federal rarity status
129 CA rarity status 1 Threatened
129 CA rarity status 2 Endangered
129 CA rarity status 3 Without CA state rarity status
130 CA nativity 1 native plants
130 CA nativity 2 alien plants
130 CA nativity 3 casual alien plants
130 CA nativity 4 naturalized plants
130 CA nativity 5 invasive plants
130 CA nativity 6 transformers
130 CA nativity 7 weeds
131 Native range 1 CA
131 Native range 2 WNA
131 Native range 3 ENA
131 Native range 4 Mexico and CAm
131 Native range 5 South America
131 Native range 6 Central Chile med region
131 Native range 6 Europe
131 Native range 7 Meditteranean Basin
131 Native range 8 Africa north of Sahara
131 Native range 9 Africa south of Sahara
131 Native range 10 South Africa cape region
131 Native range 11 Central Asia
131 Native range 12 Asia incl. temperate China
131 Native range 13 South-east Asia
131 Native range 14 Australia
131 Native range 15 Western Australia med region
131 Native range 16 Polynesia
134 CA endemicity 1 yes
134 CA endemicity 2 no
135 CFP endemicity 1 yes
135 CFP endemicity 2 no
136 endemic type 1 neoendemic
136 endemic type 2 paleoendemic

Last modified: 7/21/06, D. Ackerly