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Scleropodium touretii (Bridel) L. Koch [Brachytheciaceae]
map of distribution

Rev. Bryol. Lichenol. 18: 177. 1949. -- Hypnum touretii Brid., Musc. Recent. Suppl. 2: 185. 1812. -- Scleropodium illecebrum aucts. non B.S.G.

Plants in very dense mats, green to yellowish or olive-green. Stems to 8 cm long, prostrate, irregularly branched and markedly julaceous with branches ascending from the base but mostly curled down toward the substratum. Leaves of stems and branches similar, to 2 mm, 1.5–2.5: 1, symmetric, oblong-ovate to elliptic, widest near middle, blunt and mostly obtuse at apex but a few leaves with abrupt acumina to 0.5 mm long, concave, densely imbricated, not much changed on drying. Margins mostly minutely recurved near insertion but sometimes nearly plane, neither decurrent nor cordate at base, nearly entire below the middle but serrulate to serrate distally. Median cells smooth, to 6 µm wide, linear-vermiculate, 10–20: 1, rather thin-walled. Alar cells undifferentiated or few and quadrate to short rectangular, thin-walled, only gradually differentiated from adjacent laminal cells. Basal cells well-demarcated in 1–3 rows, rectangular. 2–3: 1, thin-walled and translucent. Differentiated basal cells cloaking base of costa. One or a group of apical cells rhomboidal, 1–3: 1. Costa extending about 3/4–7/8, usually ending in one or several abaxial spines. Pseudoparaphyllia lanceolate, sharply dentate and acuminate, to 100 µm long. Axillary hairs 4-celled, to 50 µm with 2 basal brown cells, not offset from leaf insertion. Rhizoids red-brown, sparingly branched, to 15 µm at insertion, smooth, in scattered but dense fascicles on ventral surface of stem, especially near branch insertion.
    Dioicous with male plants similar to females. Perichaetia on stoloniferous axis or on bases of branch axes, to 2.5 mm long. Perichaetial bracts ecostate, abruptly acuminate with divergent acumens from an ovate-lanceolate base; cells to 25:1, with straight lateral walls. Seta to 1.5 cm long, red-brown, smooth below but lightly papillose and geniculate at apex. Capsule brown but dark at base, strongly strangulate. Urn to 2 mm, 3–5: 1. Exothecial cells at middle of urn in more or less regular rows, to 25 µm wide, 2–3: 1, with thin but collenchymatous red-brown walls. Exothecial cells near mouth of capsule shorter and thicker walled than median exothecial cells, not in a well-demarcated band. Stomata numerous, phaneroporous, restricted to base of capsule. Operculum conic to short-rostrate, about 1/2 as long as urn. Exostome to 500 µm long, horizontally striate below, papillose above, red-brown, strongly trabeculate ventrally. Endostome segments low papillose, open on keel, about as long as exostome with basal membrane about as high as the segments. Cilia 2–3, somewhat appendiculate. Spores smooth to inconspicuously papillose, 15–18 µm.

Mail a correction to Paul Wilson ·
LiteratureBourell 1981; Flowers 1973; Harpel 1980a; Harthill et al. 1979; Holmberg 1969; Kellman 2003; Koch 1950a, 1951e, 1958; Koch and Ikenberry 1954; Lawton 1971; Long 1978; McCleary 1972; Mishler 1978; Shevock and Toren 2001; Sigal 1975; Smith 1970; Spjut 1971; Toren 1977; Whittemore and Sommers 1999; Yurky 1990, 1995. As Hypnum illecebrum Brandegee 1891; Lesquereux 1868; Watson 1880. As Scleropodium illecebrum Kingman 1912; Millspaugh and Nuttall 1923; Sayre 1940; Steere 1954.
IllustrationsMalcolm et al. 2009 p. 275; Flowers 1973; Lawton 1971; Sharp et al. 1994; Smith 1978.
BioregionsCW, NW, SN, SW.
VouchersGlenn Co.: Highway 36 at Georgie Dells Road, Norris 56061; Humboldt Co.: South Fork Mountain Road about 8 air miles north of Mad River, Six River National Forest, Norris 83883; Riverside Co.: near Ortega Oaks Campground, Norris 58148; San Mateo Co.: Spring Creek above reservoir, Filoli Center just north of Woodside, Whittemore & Noyes 4386 (UC); Tulare Co.: Marble Falls trail above Potwisha Campground, Marble Fork Kaweah River, Sequoia National Park, Shevock 12963.
   

Elevation by latitude plot for Scleropodium touretii
   in California

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