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Metaneckera menziesii (Drummond) W. C. Steere [Neckeraceae]
map of distribution
photo by Wilson of Coleman 1599drawing by Montalvo of Norris 21697

Neckera menziesii Hooker in Drummond. Musci Amer. 162. 1828.
   Neckeradelphus menziesii. (Hooker) Steere, The Bryologiat 44: 147, 1941.


    Plants in pendent streamers or in dense mats, green to yellow-green, sparingly branched. Primary stems (stolons) short and prostrate with secondary stems to 40 cm long, prostrate to pendent. Leaves to 3.5 mm, 3–6:1, asymmetric, complanate; undulate, ovate-lanceolate, acuminate to a very narrow apex, loosely imbricate when dry and little changed when moist; Margins plane, not decurrent, serrulate to serrate above the middle, sharply dentate with many recurved teeth at apex Median cells smooth, 5–8 µm wide, 5–10:1, thick-walled with lumen/wall ratio to 2–4:1, but not very pitted. Alar cells little differentiated or with a patch of thin-walled, nearly isodiametric cells in the cordate base. Median basal cells more elongate, thick-walled (lumen/wall ratio 1–2:1) and pitted. Apical cells little differentiated. Costa short and double. Pseudoparaphyllia deltoid to orbicular, strongly overarching the bud. Axillary hairs 5-celled, to 180 µm with two basal brown cells, not offset from leaf insertion. Rhizoids brown, 6–8 µm, smooth, inserted on stem immediately below abaxial costal insertion.
    Dioicous with male plants similar to females. Perigonia on main axis or terminal on short lateral branches. Perichaetia on main axis, to 6 mm long in an acicular cluster, strongly convolute around the seta. Lower perichaetial bracts ovate, abruptly contracted to acuminate, reflexed apex. Upper perichaetial bracts to 4 mm, abruptly acuminate from a long ligulate base, cells long rectangular, to 15:1. Seta 4–5 mm long, short-exserted. Capsule brown, curved and inclined, not distorted or sulcate when dry, somewhat strangulate at mouth. Urn to 2.0 mm, about 2–3:1. Exothecial cells isodiametric, hexagonal, to 35 µm wide, thin-walled, stomata restricted to neck. Operculum short-rostrate, about 1/3 as long as urn. Exostome smooth to low papillose, light-brown. Endostome segments smooth, open on keel, narrow, about as long as exostome with basal membrane less than 1/4 of total length. Cilia absent. Spores papillose, 18–20 µm.

Mail a correction to Paul Wilson ·
LiteratureBourell 1981; Holmberg 1969; Jamieson 1969; Kellman 200; Showers 1982; Spjut 1971; Steere 1967; Toren 1977; Yurky 1990, 1995. As Neckera menziesii Flowers 1973; Lawton 1971; Lesquereux 1868; Watson 1880. As Neckeradelphus menziesii Koch 1950a, 1951e; Koch and Ikenberry 1954; Steere 1941; Thomson and Ketchledge 1958.
IllustrationsMalcolm et al. 2009 p. 307; Flowers 1973; Lawton 1971.
BioregionsCaR, CW, NW, SN.
VouchersEl Dorado Co.: Sly County Park, Norris 58585; Humboldt Co.: Redwood Creek near Orick, Redwood National Park, Norris 22363; Lake Co.: Cold Creek west of Low Gap between Sheetiron Mountain and Snow Mountain, Mendocino National Forest, Shevock, Toren, & Dearing 18791; Mariposa Co.: Highway 41 near Wawona Entrance Station,Yosemite National Park, Shevock 21437; Santa Cruz Co.: Laguna Creek, Wilder Ranch State Park, Kellman 1862 (CAS); Shasta Co.: Fall Creek Road about 1 mile southeast of McCloud Bridge at Shasta Lake, Shasta-Trinity National Forest, Norris 84858; Sonoma Co.: Wolf Creek Road about 4 miles east of Gualala, Norris 86962; Tulare Co.: Lake Canyon about 0.5 mile below Oriole Lake, Sequoia National Park, Shevock 17611.
   

Elevation by latitude plot for Metaneckera menziesii
   in California

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