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Kindbergia oregana (Sullivant) Ochyra [Brachytheciaceae]
map of distribution

Plants in cushions, mats or pendent streamers, olive-green to whitish-green, often with a russet cast. Stems to 10 cm long, ascending to erect; or pendent to 50 cm long. Pinnately branched and julaceous with branches of pendent plants microphyllous and attenuate, sometimes with the entire plant composed of such branchlets. Leaves to 3.5 mm, 4–7:1, symmetric, sagittate- to ovate-lanceolate, acuminate, often with the apex twisted, loosely imbricate when dry and little changed when moist. Margins plane but sometimes ascending in the acumen, not decurrent, entire to serrulate below the middle, coarsely serrate above. Median cells mostly smooth but usually with some dorsal prorations in distal portion of leaf, 5–12 µm wide, 6–10:1, moderately thick-walled with lumen/wall ratio to 2:1, usually pitted. Alar cells quadrate and very thick-walled (lumen/wall ratio 0.5–1:1), with intensely green contents, abruptly differentiated from adjacent laminal cells, often in an excavate pocket. Apical cells rhomboidal, nearly isodiametric. Costa extending about 3/4, usually ending in an abaxial spine. Pseudoparaphyllia deltoid to orbicular, strongly overarching the bud. Axillary hairs 5-celled, to 120 µm with 2 basal brown cells, not offset from leaf insertion. Rhizoids red-brown, 10–12 µm, smooth, in scattered fascicles on ventral portion of prostrate stem.
    Dioicous with male plants similar to females. Perigonia on main axis or on larger lateral branches. Perichaetia on main axis, to 2.5 mm. Perichaetial bracts ecostate, abruptly acuminate from a ligulate-lanceolate base, cells long vermicular, to 25:1. Seta to 2 cm long, exserted. Capsule brown, inclined to horizontal, not distorted or sulcate when dry. Urn to 2 mm, 2–3:1. Exothecial cells in regular vertical rows, to 25 µm wide, 1–2:1, with thick (lumen/wall ratio 4–6:1), red-brown walls, stomata restricted to neck. Operculum conic to short-rostrate, about 1/4 as long as urn. Exostome horizontally striate below, papillose above, yellow-brown. Endostome segments low papillose, open on keel, about as long as exostome with basal membrane about as high as the segments. Cilia 2, not appendiculate. Spores papillose, 15–20 µm.

Matteri, Celina M. & Ochyra, R. 1989. Notes on two southern South American species of Brachytheciaceae (Musci). J. Hattori Bot. Lab. 66: 321–330. ( Treats of a Falkland record of K. praelonga).

Mail a correction to Paul Wilson ·
LiteratureShevock and Toren 2001; Yurky 1990. As Eurhynchium brittoniae Koch 1950a; Lawton 1971. As Eurhynchium oreganum Holmberg 1969; Howe 1897; Jamieson 1969; Kellman 2003; Koch 1950a; Koch and Ikenberry 1954; Lawton 1971; Smith 1970; Spjut 1971; Thomson and Ketchledge 1958; Yurky 1995. As Hypnum oreganum Lesquereux 1868; Sullivant 1856; Watson 1880. As Stokesiella oregana Bourell 1981; McGrew 1976; Toren 1977.
IllustrationsMalcolm et al. 2009 p. 266; Lawton 1971.
BioregionsCaR, CW, MP, NW, SN, SW.
VouchersMonterey Co.: Salmon Creek Trail, Santa Lucia Mountains, Los Padres National Forest, Norris 68223; San Francisco Co.: Mt. Davidson, Shevock 18923; Santa Barbara Co.: Windmill Canyon, Santa Rosa Island, Channel Islands National Park, Shevock 20901; Shasta Co.: Highway 299 on north side of Haynes Flat about 3 miles west of Burney, Norris 68454; Siskiyou Co.: west of White Mountain near Cook and Green Pass, Klamath National Forest, Norris 50185; Sonoma Co.: Gualala River near Lee Noble Road, Norris 86977; Tulare Co.: Freeman Creek Grove, Lloyd Meadows Basin, Sequoia National Forest, Shevock 3890.
   

Elevation by latitude plot for Kindbergia oregana
   in California

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