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Grimmia moxleyi R. S. Williams in Holzinger [Grimmiaceae]
map of distribution

Plants to 1.5 cm high, in hoary green to gray-green cushions, opaque-green to nearly black below. Leaves loosely imbricate when dry, loosely erect-spreading when moist, somewhat concave but not at all keeled, to 3 mm long with the awn constituting nearly 1/2 of that length, ovate-lanceolate from a loosely sheathing somewhat expanded base, 2.5–3.5: 1. Median leaf cells in regular longitudinal rows, unistratose in the expanded base but uniformly bistratose in the limb, smooth, to 7 µm wide, isodiametric with lumens irregular but without corner thickenings. Marginal cells not differentiated. Basal juxtacostal cells elliptical, to 3: 1, to 15 µm wide, with irregularly thickened lateral walls. Basal marginal cells transversely elongate, 0.5–0.7: 1, to 15 µm wide. Costa flared at the base and filling 1/3–1/4 of immediate leaf base, narrowed and obscure above, percurrent in the somewhat acuminate awn. Costa cross-section at leaf middle broad and flattened, less than 1.5 times as thick as the adjacent leaf, with a weak abaxial band of stereids or substereids and with a single layer of larger cells on the adaxial face. Margin of leaf limb plane to incurved, entire below. Awn strongly decurrent with its margin somewht serrulate, and with its surface spinose-prorate. Axillary hairs to 100 µm long, with 6–8 nearly isodiametric and thick- walled cells, without a basal brown cell. Rhizoids red brown, to 15 µm wide at base, smooth. Stem cross-section rounded-triangular with a large and well-defined central strand, and with leptodermous, hyaline and large inner cortical cells, and with the 1–2 cell-thick outer stereome composed of much smaller, thick-walled cells strongly pigment with yellow to yellow-brown.
    Dioicous with male and female plants similar. Perichaetial leaves loosely sheathing base of seta, somewhat larger and with a relatively longer, less decurrent awn than the adjacent vegetative leaves. Seta to 2.5 mm long, yellow to pale brown, straight and smooth. Urn erect, symmetrical, smooth to lightly wrinkled when dry, to 1.2 mm long, 1.5–2: 1, not strumose. Operculum short rostrate, about 1/3 as long as urn. Annulus formed of 2–3 rows of relatively more thin-walled cells. Calyptra mitrate, covering about 1/2 of urn. Peristome red-brown, to 200 µm long, cribrose along median line near the apex, papillose to vertically striate-papillose. Exothecial cells to 25 µm wide, not in regular vertical rows, quite irregular in size and shape, rectangular 2–4: 1, rather thick walled. Stomata phaneroporous, restricted to base of urn. Spores smooth, to 12 µm.

Mail a correction to Paul Wilson ·
LiteratureGreven 2003; Harthill et al. 1979; Koch 1950a; Long 1978; Moxley 1928.
IllustrationsGreven 2003.
NotesAccording to Muñoz (pers. comm.), Grimmia moxleyi represents the long hair point expression of the widespread taxon Grimmia orbicularis. Roxanne Hastings (pers. comm.) and Greven (2003), however, recognize this taxon at the species level and this is how it will be treated in the Flora of North America. Regardless of circumscription, this ecological expression is common in the Mojave Desert and easily recognized in the field by the exceedingly long white hairpoints.
BioregionsDMoj, DSon.
VouchersAngeles Co.: Big Rock Creek and Devils Punchbowl, Angeles National Forest, Moxley 1141 (NY, MO) and Tick Canyon, MacFadden 19695 (CAS); Inyo Co.: Fall Canyon just north of Titus Canyon, Grapevine Mountains, Death Valley National Park, Shevock, Harpel, & York 19045.
   

Elevation by latitude plot for Grimmia moxleyi
   in California

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