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Grimmia anomala Hampe in W. P. Schimper [Grimmiaceae]
map of distribution

Syn. ed. 2: 270. 1876. -- Grimmia hartmanii var. anomala (Hampe) Moenk., Laubm. Eur. 369. 1927. -- Dryptodon anomalus (Hampe) Loeske, Hedwigia 49: 32. 1909. -- Grimmia elatior ssp. anomala (Hampe) Kindb., Eur. & N. Am. Bryin. 222. 1897.
   Grimmia philibertiana Britt., Bull. Torr. Bot. Club 18: 51. 1891.

Grimmia anomala Hampe ex W.P.Schimper, Syn. Musc. Eur. ed. 2: 270.
   
    Grimmia elatior ssp. anomala (Hampe) Kindberg. Eur. N. Am.Bryin 2:222. 1899
    Grimmia hartmanii var. alpinoborealis Loeske. Laubm, Eur. Part 1. : 141. 1913.

Dryptodon anomalus (Hampe) Loeske. Hedwigia 49: 32. 1909
    Grimmia hartmanii var. anomala (Hampe) Moenkemeyer. Laubm Eur.369. 1932.
    Grimmia hartmanii ssp. anomala (Hampe ) Loeske. Biblioth. Bot. 101: 182. 1930.

Plants to 1.5 cm high, in hoary green to gray-green cushions, opaque-green to nearly black below. Leaves loosely imbricate when dry, loosely erect-spreading when moist, somewhat concave but not at all keeled, to 3 mm long with the awn constituting nearly 1/2 of that length, ovate-lanceolate from a loosely sheathing somewhat expanded base, 2.5–3.5: 1. Median leaf cells in regular longitudinal rows, unistratose in the expanded base but uniformly bistratose in the limb, smooth, to 7 µm wide, isodiametric with lumens irregular but without corner thickenings. Marginal cells not differentiated. Basal juxtacostal cells elliptical, to 3: 1, to 15 µm wide, with irregularly thickened lateral walls. Basal marginal cells transversely elongate, 0.5–0.7: 1, to 15 µm wide. Costa flared at the base and filling 1/3–1/4 of immediate leaf base, narrowed and obscure above, percurrent in the somewhat acuminate awn. Costa cross-section at leaf middle broad and flattened, less than 1.5 times as thick as the adjacent leaf, with a weak abaxial band of stereids or substereids and with a single layer of larger cells on the adaxial face. Margin of leaf limb plane to incurved, entire below. Awn strongly decurrent with its margin somewht serrulate, and with its surface spinose-prorate. Axillary hairs to 100 µm long, with 6–8 nearly isodiametric and thick- walled cells, without a basal brown cell. Rhizoids red brown, to 15 µm wide at base, smooth. Stem cross-section rounded-triangular with a large and well-defined central strand, and with leptodermous, hyaline and large inner cortical cells, and with the 1–2 cell-thick outer stereome composed of much smaller, thick-walled cells strongly pigment with yellow to yellow-brown.
    Dioicous with male and female plants similar. Perichaetial leaves loosely sheathing base of seta, somewhat larger and with a relatively longer, less decurrent awn than the adjacent vegetative leaves. Seta to 2.5 mm long, yellow to pale brown, straight and smooth. Urn erect, symmetrical, smooth to lightly wrinkled when dry, to 1.2 mm long, 1.5–2: 1, not strumose. Operculum short rostrate, about 1/3 as long as urn. Annulus formed of 1 row of relatively more thin-walled cells. Calyptra mitrate, covering about 1/2 of urn. Peristome red-brown, to 200 µm long, cribrose along median line near the apex, papillose to vertically striate-papillose. Exothecial cells to 25 µm wide, not in regular vertical rows, quite irregular in size and shape, rectangular 2–4: 1, rather thick walled. Stomata phaneroporous, restricted to base of urn. Spores smooth, to 12 µm.

Mail a correction to Paul Wilson ·
LiteratureMisapplied as Grimmia hartmannii Harpel 1980a; Harthill et al. 1979; Koch 1950a; Lawton 1971; Long 1978; Mishler 1978; Showers 1982; Spjut 1971.
IllustrationsMalcolm et al. 2009 p. 62; Greven 2003; Ignatov and Ignatova 2003. As Grimmia hartmannii var. anomala Crum and Anderson 1981; Flowers 1973.
BioregionsCaR, NW.
VouchersDel Norte Co.: Doctor Rock, Norris 57676 (determined by Muñoz); El Dorado Co.: Ropi Lake, Norris 71109 (determined by Muñoz); Humboldt Co.: Big Hill Road at South Fork Mill Creek, Six Rivers National Forest, Norris 58636; Plumas Co.: Quincy-La Porte Road at Onion Valley, Plumas National Forest, Norris 83187; Tehama Co.: Doll Basin Research Natural Area, Mendocino National Forest, Toren & Bourell 6666 (CAS).
   

Elevation by latitude plot for Grimmia anomala
   in California

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