Plants light green to rather deep green, to 6 mm high, with up to 10 pairs of leaves. Leaves, when dry, laterally twisted; when moist, spreading from stem at about a 30 degree angle, to 2 mm long, 3–4: 1 oblong to oblong lingulate. Vaginant lamina to 5:1, extending about 1/2 of leaf, the two sides converging at an acute angle, homostichous. Dorsal lamina mostly tapered to extinction above the leaf base, not decurrent. Median cells of apical and dorsal laminae unistratose, smooth, to 12 µm wide, thin-walled, hexagonal to rhomboidal with angular lumens and no corner thickenings. Median cells of vaginant lamina somewhat more regularly arranged, mostly a little longer than wide, 1–2:1, to 12 µm wide. Basal juxtacostal cells somewhat larger and more elongate. Costa to 25 µm at base, percurrent to subpercurrent, sometimes merging with limbidium at apex and forming an apiculus. Margin plane and entire throughout. Limbidium variably present, extending to apex or ending several cells below apex, occasionally almost absent from dorsal lamina. up to 4 cells wide, without stereids, not at all decurrent. Stem with no central strand and with a few thin-walled inner cortical cells abruptly differentiated from 1–2 layers of smaller, rather thick-walled outer corticals; lateral cells of outer cortex poorly differentiated. Axillary hyaline nodules not present.
Dioicous or autoicous. Perichaetia terminal with seta to 8 mm, red-brown, nearly straight above the geniculate base, smooth. Urn to 0.8 mm long, oblong and symmetrical, about 2: 1. Operculum conic, about 1/3 as long as urn. Calyptra cucullate, smooth. Peristome reddish-brown, to 250 µm long, minutely papillose proximally, spirally ridged distally. Exothecial cells in regular vertical rows, somewhat bulging on outer surface, mostly isodiametric with angular lumens, rather thin-walled, to 15 µm wide. Spores to 20 µm, finely papillose.
|Literature||Kellman 2003; Pursell 1997a; Shevock and Toren 2001.|
|Illustrations||Pursell 1997a; Sharp et al. 1994.|
|Bioregions||CaR, CW, DMoj, DSon, MP, NW, SN, SNE, SW.|
|Vouchers||Kern Co.: Short Canyon Trail west of L.A. Aqueduct, Owens Peak BLM Wilderness, Shevock & Newberry 16979 (determined by Pursell); Lake Co.: Hidden Valley north of Middletown, Toren & Dearing 7373a (CAS) [determined by Pursell]; Lassen Co.: Johnsondale Dump Road southeast of Susanville, Norris 52838; Los Angeles Co.: La Tuna Canyon, Verdugo Hills, MacFadden 8085 (MO) [determined by Pursell]; San Bernardino Co.: Keystone Canyon, New York Mountains, Mojave National Preserve, Laeger & Davis 1328 (CAS, PAC) [determined by Pursell]; San Diego Co.: Highway 94, 2 miles west of Tecate Road, Stark 813 (MO) [determined by Pursell]; San Francisco Co.: Coastal Bluff Trail, Presidio of San Francisco, Shevock 18945 (determined by Pursell); Santa Barbara Co.: Water Canyon, Santa Rosa Island, Channel Islands National Park, Shevock & Rodriquez 20832; Shasta Co.: Highway 299 about 9 miles east of Bella Vista, Norris 47616.|