The Jepson Manual: Higher Plants of California

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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

Names are linked to treatments from the Manual


Key to BOTRYCHIUM

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Sterile leaf part generally > 10 cm; leaf bud densely hairy; fertile leaf part aborted or well developed; leaf leathery, evergreen, each part generally ± 2–3-pinnate
B. multifidum
  1' Sterile leaf part generally < 10 cm; leaf bud glabrous; fertile leaf part well developed; leaf herbaceous, deciduous, each part simple to ± 2-pinnate
 
    2. Sterile leaf part segments oblique-ovate to lanceolate-oblong, midribbed
B. pinnatum
    2' Sterile leaf part segments generally spoon- to wedge-or fan-shaped, not midribbed
 
      3. Sterile and fertile leaf parts separated near to well below mid-leaf; blade tips coarsely lobed to irregularly angled or shallowly rounded
 
        4. Sterile, fertile leaf parts separated near to just below middle of leaf; full-sized blades oblong, non-ternate, segments irregularly angled, square or oblong; shady coniferous woods
B. montanum
        4' Sterile, fertile leaf parts separated well below middle of leaf, generally at top of leaf sheath; full-sized blades oblong to ovate, ternate, segments wedge- to fan-shaped; open marshes, damp meadows
B. simplex
      3' Sterile and fertile leaf parts separated near or generally above mid-leaf; blade tips finely, regularly, and deeply lobed
 
        5. Lower segments of sterile leaf part widely fan-shaped, margins at base meeting at 120–160°
 
          6. Plant slender, herbaceous-fleshy; sterile leaf part generally < 6 cm, < 2 cm wide, segments 3–5 pairs, ± well separated, outer margins generally finely crenate to dentate; fertile leaf part 1.3–3 × sterile leaf part; marshes, meadows
B. crenulatum
          6' Plant robust, fleshy; sterile leaf part generally 6–10 cm, 2–4 cm wide, segments 4–9 pairs, generally touching to overlapped, outer margins generally entire; fertile leaf part 0.8–2 × sterile; dryish fields, meadows
B. lunaria
        5' Lower segments of sterile leaf part narrowly fan- to wedge-shaped, margins at base meeting at 40–100(120)°
 
          7. Segments of sterile leaf parts distally angled, irregularly toothed, ascending, lower segments wedge-shaped, margins at base meeting at 40–90(120)°, basal often with scattered marginal sporangia
B. ascendens
          7' Segments of sterile leaf parts distally rounded, ± smooth, not or ± ascending, lower segments ± fan-shaped, margins at base meeting at 50–100(120)°, basal without scattered marginal sporangia
B. minganense


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