The Jepson Manual: Higher Plants of California

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Names are linked to treatments from the Manual


Key to VIOLA

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Stipule leaf-like, ± = leaf, pinnately lobed; generally annual; alien
 
    2. Petals generally < sepals, mostly cream, sometimes with some blue-violet on upper 4, spur ± = sepal appendages
V. arvensis
    2' Petals generally > sepals, mainly blue-violet (rarely yellow), spur > sepal appendages — sometimes per
V. tricolor
  1' Stipule not leaf-like, << leaf, entire or toothed; perennial herb; generally native
 
    3. Leaf dissected to compound
 
      4. Cleistogamous flowers present, sometimes exclusively; leaf blade generally wider than long
V. sheltonii
      4' Cleistogamous flowers 0; leaf blade generally longer than wide
 
        5. Upper 2 petals golden-yellow on front, brown to very dark on back
V. douglasii
        5' Upper 2 petals dark red-violet
 
          6. Lower 3 petals bluish to white, bases with yellow area; stem glabrous to puberulent
V. beckwithii
          6' Lower 3 petals cream, bases with deep yellow area; stem glabrous
V. hallii
    3' Leaf generally ± entire to deeply lobed (seldom dissected)
 
      7. Petals white to blue or purple, without yellow
 
        8. Stem branched; leaves basal and cauline
V. adunca
        8' Stem 0; leaves basal
 
          9. Petals white, dark-veined or not
 
            10. Leaf blade elliptic to ovate, longer than wide — nw Klamath Ranges, 100–500 m
V. primulifolia subsp. occidentalis
            10' Leaf blade ovate to reniform, generally as wide as or wider than long
 
              11. Rhizome < 5 mm diam; spur white; leaf blade thin, not leathery, (sub)entire; native; mtns
V. macloskeyi
              11' Rhizome 5–20 mm diam; spur violet; leaf blade thick, leathery, ± toothed; alien; San Francisco Bay Area, South Coast
2 V. odorata
          9' Petals blue (sometimes very pale) to purple
 
            12. Leaf glabrous; rhizome thin; North Coast
V. palustris
            12' Leaf ± glabrous or finely hairy; rhizome thick; Klamath Ranges, San Francisco Bay Area, South Coast, San Bernardino Mountains, Warner Mountains, East of Sierra Nevada
 
              13. Plant stoloned; leaf finely hairy; alien; San Francisco Bay Area, South Coast
2 V. odorata
              13' Plant not stoloned; leaf ± glabrous; native; Klamath Ranges, San Bernardino Mountains, Warner Mountains, East of Sierra Nevada
V. sororia subsp. affinis
      7' Petals yellow to ± orange, or white with yellow base and spur
 
        14. Petals white inside, with generally yellow base and spur, lower 3 veined purple-red
 
          15. Leaf blade base generally tapered; basal blade midvein 10–40 mm; plant glabrous
V. cuneata
          15' Leaf blade base generally truncate to deeply cordate; basal blade midvein 10–60 mm; plant finely hairy
V. ocellata
        14' Petals yellow to ± orange inside, generally veined dark brown to purple
 
          16. Stems few, erect, unbranched below; basal leaves 0–4; cauline leaves and peduncles restricted to upper stem
 
            17. Upper 2 petals yellow outside; cauline leaf ovate-cordate to widely reniform; moist places
V. glabella
            17' Upper 2 petals purple to brown-purple outside; cauline leaf diamond-shaped to reniform, entire to dissected; dry places
V. lobata
              18. Cauline leaf diamond-shaped to generally deltate, entire to toothed
subsp. integrifolia
              18' Cauline leaf ± ovate to generally reniform, palmately lobed or dissected, segments 3–12, generally entire
subsp. lobata
          16' Stems 0–many, prostrate to erect, branched below; basal leaves 0–6; cauline leaves and peduncles on lower and upper stem
 
            19. Plant stoloned; leaf evergreen, leathery, generally purple-spotted on 1 or both sides — coastal forests
V. sempervirens
            19' Plant not stoloned; leaf not evergreen, not leathery, not purple-spotted
 
              20. Basal leaves 0; cleistogamous flowers 0; petals orange-yellow — fruit glabrous
V. pedunculata
              20' Basal leaves 1–6; cleistogamous flowers present; petals yellow to lemon-yellow
 
                21. Basal blade 1.5–6 × longer than wide, 15–80 mm, linear to obovate; cauline blade 2–10 × longer than wide
 
                  22. Plant white-tomentose — n&c High Sierra Nevada
V. tomentosa
                  22' Plant glabrous to densely hairy (not white-tomentose)
 
                    23. Leaf generally entire
V. bakeri
                    23' Leaf toothed, jagged, or wavy (seldom entire)
 
                      24. Cauline leaf blade 3–25 mm wide, linear to ± ovate; peduncle 20–90 mm
V. pinetorum
                        25. Plant canescent, 4–10 cm; leaves 3–10 mm wide, basal 40–95 mm, cauline 30–80 mm — s High Sierra Nevada
subsp. grisea
                        25' Plant glabrous to hairy, 6–22 cm; leaves 4–25 mm wide, basal 50–200 mm, cauline 30–150 mm
subsp. pinetorum
                      24' Cauline leaf blade 14–45 mm wide, elliptic to ± ovate; peduncle 40–180 mm
V. praemorsa
                        26. Cauline leaf blade much longer than wide, generally irregularly toothed or wavy, base generally tapered
subsp. linguifolia
                        26' Cauline leaf blade slightly longer than wide, generally regularly toothed, base ± truncate to subcordate
subsp. praemorsa
                21' Basal blade generally 1–1.5 × longer than wide, 10–50 mm, ovate to round; cauline blade generally < 2.5 × longer than wide
 
                  27. Leaf canescent above; plant generally 6–13 cm; peduncle 30–100 mm; uncommon — East of Sierra Nevada, Mojave Desert
V. aurea
                  27' Leaf glabrous to hairy (not canescent) above; plant 1–35 cm; peduncle 15–170 mm; widespread
V. purpurea
                    28. Plant 1–12 cm; stem generally buried, little branched or elongated by late summer; lower petal (including spur) generally < 11 mm
 
                      29. Cauline leaf generally entire; plant 1–12 cm
subsp. integrifolia
                      29' Cauline leaf generally crenate to deeply toothed; plant 4–11 cm
subsp. venosa
                    28' Plant 4–35 cm; stem generally not buried, clearly branched and elongated above by late summer; lower petal (including spur) generally > 11 mm
 
                      30. Cauline leaf deeply toothed
subsp. mohavensis
                      30' Cauline leaf entire, jagged, shallowly toothed, or lobed
 
                        31. Leaf generally green tinged purple, generally glabrous above; lower petal (including spur) generally < 13 mm; peduncle generally < 13 cm; 500–2500 m, generally montane forest
subsp. purpurea
                        31' Leaf generally gray-green (± without purple), puberulent; lower petal (including spur) generally 13+ mm, or peduncle > 13 cm, or both; 400–1800 m, generally foothill communities
subsp. quercetorum


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