The Jepson Manual: Higher Plants of California

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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

Names are linked to treatments from the Manual


Key to VERBENA

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Corolla 8–17 mm, limb 6–10 mm wide; calyx 5–9.5 mm
 
    2. Leaves 3–5-lobed; calyx hairs spreading
V. gooddingii
    2' Leaves 1–2-pinnately dissected; calyx hairs appressed
V. tenuisecta
  1' Corolla 1.5–6 mm, limb < 5 mm wide; calyx 2.5–4 mm
 
    3. Flower bract 4–8 mm, 2–3 mm > calyx; most stems prostrate to decumbent
V. bracteata
    3' Flower bract 1–4.5 mm, generally < calyx; most stems ascending to erect
 
      4. Most cauline leaf blades 1–2-lobed (lobes below middle 8–25 mm deep), toothed
 
        5. Herbage sparsely strigose; calyx minutely strigose
V. menthifolia
        5' Herbage and calyx short-spreading-(generally soft-)hairy
2 V. lasiostachys
          6. Leaf surface grayish green above, hairs generally soft; nutlet scar brownish at 20×
var. lasiostachys
          6' Leaf surface green above, hairs generally stiff; nutlet scar white-papillate at 20×
var. scabrida
      4' Most cauline leaf blades generally unlobed (except some V. hastata), teeth 2–8 mm
 
        7. Lower and mid-cauline petioles generally 1–2.5 cm, sometimes narrowly winged
 
          8. Petioles, inflorescence axis, and calyx short-spreading-(generally soft-) hairy
2 V. lasiostachys (see 6. for vars.)
          8' Petioles, inflorescence axis, and calyx strigose or scabrous
 
            9. Fruits generally overlapping (axis with 8–10 fruits per cm); upper surface of leaves rough-puberulent to slightly scabrous
V. hastata
            9' Fruits not overlapping, especially below (axis generally with < 8 fruits per cm); upper surface of leaves strigose to scabrous
V. scabra
        7' Lower and mid-cauline petiole generally 0 (blade often clasping)
 
          10. Fruits not overlapping (especially below middle); flower bracts 3.5–4 mm; calyx 4–4.5 mm; spikes 1–5 per cluster
V. californica
          10' Fruits overlapping; flower bracts 3–3.5 mm; calyx 3–3.5 mm; spikes generally 5–17 per cluster
 
            11. Leaf base cordate to truncate, generally clasping; spikes ± 5–6 mm diam in fruit
V. bonariensis
            11' Leaf base ± acute, subsessile; spikes 3–4 mm diam in fruit
V. litoralis


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