| 1. Herbage green, ± sticky-glandular, short-hairy |
| O. tauricum |
| 1' Herbage ± canescent-tomentose |
| |
| 2. Leaves dentate to shallowly lobed; phyllaries linear; corolla lobes not glandular |
| O. acanthium subsp. acanthium |
| 2' Leaves ± deeply 12 × lobed; phyllaries lanceolate to ovate; corolla lobes glandular-puberulent |
| O. illyricum
|