The Jepson Manual: Higher Plants of California

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Names are linked to treatments from the Manual


Key to LOTUS Key to Groups

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Leaflets 5, 3 near axis tip, lower pair large, stipular in position; stipules gland-like, often not apparent
Group 1
  1' Leaflets 3–15, lowest not stipular in position; stipules as above or not
 
    2. Stipules scarious or leaflet-like, sometimes fragile and quickly lost; leaflets 3–15, pinnately arranged, often opposite or subopposite
Group 2
    2' Stipules gland-like, bump-like, or conic, often not apparent; leaflets 3–9, pinnately or ± palmately arranged, generally irregularly arrayed on leaf axis
 
      3. Fruit indehiscent, generally not flat, generally included or moderately exserted from calyx (sometimes conspicuously exserted and curved most of length), generally with curved, 2–3 mm beak; generally per
Group 3
      3' Fruit dehiscent, generally flat, generally almost entirely exserted and straight, generally with straight or curved, 0.5–1.5 mm beak; annual or perennial herb
 
        4. Annual; fruit generally straight; wings < or ± = keel; stigma glabrous
Group 4
        4' Perennial (except L. strigosus); fruit straight or ± curved; wings generally > keel; stigma nearly glabrous or finely hairy
Group 5

Key to LOTUS Group 1: Leaflets 5, lower pair stipular in position

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Annual; hairy; inflorescence generally 1–3-flowered, 5–8 mm — North Coast (Sonoma Co.)
L. angustissimus
  1' Perennial; glabrous or hairy; inflorescence 3–12-flowered, 8–14 mm
 
    2. Inflorescence 3–8-flowered; calyx lobes not outcurved in bud; rhizome 0; stem generally solid, glabrous or strigose
L. corniculatus
    2' Inflorescence 8–12-flowered; calyx lobes often outcurved in bud; rhizome conspicuous; stem hollow, hairs often spreading
L. uliginosus

Key to LOTUS Group 2: Stipules scarious or leaflet-like

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Leaflets 3–5; flower 8–10 mm; plant very small, decumbent; North Coast Ranges
L. yollabolliensis
  1' Leaflets (3)5–15; flower generally 8–15 mm; plant various, generally erect or ascending; if leaflets < 5, not of North Coast Ranges
 
    2. Leaf conspicuously hairy, gray or silvery, with soft, wavy hairs; fruit glabrous
L. incanus
    2' Leaf glabrous or hairy, rarely gray or silvery; fruit generally hairy if leaf hairy
 
      3. Peduncle bract 0 or just below umbel, generally simple or 2–3-parted; fruit 1.5–3 mm wide
 
        4. Petal claws included to slightly exserted from calyx; plant often hairy
L. oblongifolius
          5. Leaflets 3–7, glabrous; corolla 8–9 mm; local, Tulare Co.
var. cupreus
          5' Leaflets 7–11, glabrous or hairy; corolla 9–13 mm; more widespread, including Tulare Co.
var. oblongifolius
        4' Petal claws clearly exserted from calyx; plant ± glabrous
 
          6. Corolla wings pink or purple, fading white; bracts generally present and 3-parted
L. formosissimus
          6' Corolla wings white; bracts 0 or generally simple
L. pinnatus
      3' Peduncle bract generally well below umbel, generally 3–5-parted, sometimes fragile and lost; fruit 3–5 mm wide
 
        7. Hairs generally persistent, spreading, sometimes glandular; stipules often wide, leaflet-like, persistent and sometimes becoming scarious; corolla pink to red-purple; n&c California
L. stipularis
          8. Stem wiry; stipules wide, clasping stem
var. ottleyi
          8' Stem often fleshy; stipules narrow or wide, not or slightly clasping stem
var. stipularis
        7' Hairs generally nearly 0 or appressed, rarely glandular; stipules scarious, fragile, not leaflet-like, generally some lost; corolla initially white, pink, or yellow-green
 
          9. Flowers generally 6–10; corolla 10–12 mm, white or pink, becoming striate, darker; pedicel 2–3 mm, slightly longer in fruit; leaflet length generally 2–3 × width; generally n Outer North Coast Ranges
L. aboriginus
          9' Flowers generally 12–20; corolla 12–17 mm, yellow-green, becoming dark-blotched; pedicel generally 3–6 mm (> 3 mm at least in fruit); leaflet length generally 1–2.5 × width; widespread, especially inland
L. crassifolius
            10. Plant glabrous or strigose; widespread, including n Peninsular Ranges (Riverside Co.)
var. crassifolius
            10' Plant soft-hairy; s Peninsular Ranges (San Diego Co.)
var. otayensis

Key to LOTUS Group 3: Stipules often gland-like; fr indehiscent

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Annual, prostrate or decumbent, sometimes ascending; young growth not conspicuously hairy; flower 3–7 mm; fruit bodies exserted; s South Coast Ranges, Southwestern California, generally s of Los Angeles
 
    2. Inflorescence ± sessile; s South Coast Ranges, Southwestern California
L. hamatus
    2' Inflorescence conspicuously peduncled; s South Coast (San Diego Co.)
L. nuttallianus
  1' Perennial or shrub, prostrate to erect; young growth conspicuously hairy if plant blooming 1st year or flowers < 7 mm; fruit bodies included or exserted; widespread, including Southwestern California
 
    3. Perennial or shrub; Channel Islands
 
      4. Calyx nearly glabrous or strigose; fruit body soon exserted; leaf green or gray, sparsely or densely strigose or silky
L. dendroideus
        5. Leaflets 3, densely strigose or silky, generally gray; n Channel Islands (San Miguel Island)
var. veatchii
        5' Leaflets 3–5, finely or sparsely strigose, green; Channel Islands (except San Miguel Island)
 
          6. Fruit 1–1.5 cm; peduncle bract 0; Channel Islands (except San Miguel, San Clemente islands)
var. dendroideus
          6' Fruit 2.5–5 cm; peduncle bract generally present; s Channel Islands (San Clemente Island)
var. traskiae
      4' Calyx shaggy-hairy; fruit including (except slowly elongating beak); leaf generally silvery-silky with fine, straight, ultimately wavy or tangled hairs
2 L. argophyllus
        7. Inflorescences peduncled, not crowded; calyx lobes 1.5–2.5 mm; stem decumbent to ascending
var. argenteus
        7' Inflorescences ± sessile, densely crowded at stem tip; calyx lobes 2.5–5 mm; stem ascending to erect
 
          8. Stem erect; leaves overlapping; calyx lobes ± 2.5–3.5 mm; s Channel Islands (San Clemente Island)
var. adsurgens
          8' Stem ascending to erect; leaves not overlapping; calyx lobes 2.5–5 mm; n Channel Islands (Santa Cruz Island)
var. niveus
    3' Perennial, rarely shrub (except L. scoparius); mainland
 
      9. Leaflets 3; inflorescence 1–3-flowered, ± sessile
 
        10. Leaflet 2–5 mm; fruit 6–9 mm; uncommon, se Peninsular Ranges, sw Sonoran Desert
L. haydonii
        10' Leaflet 4–12 mm; fruit 10–15 mm; widespread, including Peninsular Ranges
 
          11. Calyx glabrous (see couplet 15 for vars.)
2 L. scoparius
          11' Calyx strigose
L. procumbens
            12. Corolla 9–12 mm; calyx 4–6 mm, lobes ± = tube; s Sierra Nevada
var. jepsonii
            12' Corolla 6–8 mm; calyx 2–3 mm, lobes << tube; not s Sierra Nevada
var. procumbens
      9' Leaflets > 3 on some or most leaves; inflorescence 2–15-flowered, sessile or peduncled
 
        13. Leaflets glabrous or finely strigose, generally green; hairs straight or slightly wavy
 
          14. Stem generally ascending or erect (some of immediate coast prostrate), bushy-branched, often ± 1 m; stem slightly woody (but green), sparsely leafy; corolla yellow; calyx lobes generally 1–2 mm, not curved outward; coastal and inland
2 L. scoparius
            15. Keel prominent, > wings; flower generally 8–9 mm; generally inland, Southwestern California, Sonoran Desert
var. brevialatus
            15' Keel not prominent, ± = wings; flower 7–12 mm; coastal and inland, widespread
var. scoparius
          14' Plants prostrate or low-ascending, < 1 m; stem not woody, leafy; corolla pink-white or yellow; calyx lobes generally < 1 mm or curved outward; especially coastal
 
            16. Calyx lobes slender, some or all curved outward or hooked; corolla generally white to pinkish (to brick-red when dry)
L. benthamii
            16' Calyx lobes wide, not curved outward, not hooked; corolla yellow
L. junceus
              17. Peduncle 8–25 mm; stem wiry, generally decumbent; fruit body well exserted
var. biolettii
              17' Peduncle 1–5 mm; stem generally stout, prostrate to ascending; fruit body slightly exserted
var. junceus
        13' Leaflets conspicuously hairy, green, gray, or silvery; hairs straight or wavy
 
          18. Stem, especially near tip, generally with conspicuous, spreading or obliquely directed, often straight and stiff hairs < 0.5 mm; corolla 4–6 mm; especially coastal California, inland to Desert edge
L. heermannii
            19. Corolla generally 4–5 mm; ovary, young fruit generally finely strigose; South Coast, Peninsular Ranges, Desert edge
var. heermannii
            19' Corolla generally 5–6 mm; ovary, young fruit generally with spreading hairs; North Coast, Outer North Coast Ranges, Central Coast, Outer South Coast Ranges
var. orbicularis
          18' Stem generally strigose or with ± spreading, often wavy hairs generally 0.2–0.4 mm; corolla 5–12 mm; especially inland California
 
            20. Fruit body exserted; leaflet hairs generally wavy, not obscuring leaflet surface; leaves gray or green
L. nevadensis
              21. Flowers generally 3–5 per inflorescence, 6–7 mm, appearing blocky; banner abruptly upcurved 90°; Transverse Ranges, ne Peninsular Ranges
var. davidsonii
              21' Flowers generally 5–12 per inflorescence, 5–10 mm, not appearing blocky; banner upcurved 30–90°; not of Transverse Ranges
var. nevadensis
            20' Fruit body included or shortly exserted; leaflet hairs at first generally straight, in age tangled and generally obscuring leaflet surface; leaves silky-silvery
2 L. argophyllus
              22. Inflorescence generally < 1 cm wide, 4–8-flowered; corolla 6–10 mm; especially c Sierra Nevada and s
var. argophyllus
              22' Inflorescence often 1.5 cm wide, 10–15-flowered; corolla 8–12 mm; n Sierra Nevada
var. fremontii

Key to LOTUS Group 4: Stipules often gland-like; fr dehiscent; annuals

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Flowers generally 1 per leaf axil; peduncle << 1 cm, bracts 0
 
    2. Corolla yellow, reddening with age; plant prostrate to low-ascending and calyx lobes 0.8–2 × tube
 
      3. Calyx lobes 1–2 × tube; fruit generally 3–4 mm wide; plant generally with soft, spreading hairs
L. humistratus
      3' Calyx lobes ± 0.8–1.2 × tube; fruit generally 2.3–3 mm wide; plant inconspicuously strigose or with soft, spreading hairs
L. wrangelianus
    2' Corolla white, pale yellow, or pink, often darkening with age; plant generally ascending to erect and calyx lobes < 2 × tube, or rarely prostrate and calyx lobes > 2 × tube
 
      4. Plant generally 1–3 dm; calyx lobes ± 1.5 × tube; common
L. denticulatus
      4' Plant < 1 dm; calyx lobes ± 2 × tube; uncommon
L. rubriflorus
  1' Flowers 1–several per leaf axil; peduncle generally > 1 cm, generally bracted
 
    5. Inflorescence generally 2–4-flowered (1st-formed sometimes 1–2-flowered); leaflets 3–7, obovate or ± round, terminal generally largest
L. salsuginosus
      6. Corolla 3.5–5 mm, keel > other petals; fruit 1–1.5 cm, becoming narrowed between seeds
var. brevivexillus
      6' Corolla 6–10 mm, keel ± = other petals; fruit 1.5–3 cm, not narrowed between seeds
var. salsuginosus
    5' Inflorescence generally 1-flowered; leaflets 1–5, lanceolate, elliptic, or obovate, ± equal
 
      7. Calyx lobes >> tube; leaflets generally 3 (often 1 on upper leaves)
L. purshianus
      7' Calyx lobes ± = tube; leaflets 3–9 throughout
 
        8. Stem erect or ascending; leaf axis not conspicuously flattened; corolla pink or salmon, quickly fading; fruit straight, margin often wavy
L. micranthus
        8' Stem prostrate, often mat-forming; leaf axis conspicuously flattened; corolla yellow; fruit curved near tip, margin not wavy
L. strigosus

Key to LOTUS Group 5: Stipules often gland-like; fr dehiscent; perennials

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Annual, generally prostrate and mat-forming; inflorescence generally 1–2-flowered; corolla 5–10 mm; leaf axis flat, ± blade-like; fruit generally curved only at or near tip
L. strigosus
  1' Perennial or shrub-like, prostrate (mat-forming or not) to erect; inflorescence 1-9-flowered; corolla 8–25 mm; leaf axis not flat or blade-like; fruit generally straight (sometimes curved throughout, rarely only at or near tip)
 
    2. Leaf axis including petiole 10–35 mm; leaflets 7–9, generally 1–2 cm, length 1.5–3 × width
L. grandiflorus
      3. Plant puberulent or soft-hairy; leaflet generally 1–1.5 cm
var. grandiflorus
      3' Plant ± glabrous or puberulent; leaflet generally 1.6–2.2 cm
var. macranthus
    2' Leaf axis including petiole 1–8 mm; leaflets 3–5, < 1 cm or length > 3 × width, or both
 
      4. Shrub-like, ascending, < 1.5 m, finely strigose; corolla 12–22 mm
L. rigidus
      4' Perennial, prostrate or low-ascending, < 0.3 m, silvery-silky or gray-puberulent; corolla 8–12 mm
L. argyraeus
        5. Stem prostrate, mat-forming; calyx lobes ± 2 mm; San Bernardino Mountains, San Jacinto Mountains, Desert Mountains
var. argyraeus
        5' Stem decumbent to low-ascending; calyx lobes 2–3 mm; Desert Mountains
 
          6. Leaflet oblanceolate to obovate, length ± 3–4 × width; New York Mtns
var. multicaulis
          6' Leaflet obovate, length ± 2 × width; Providence Mtns
var. notitius


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