The Jepson Manual: Higher Plants of California

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Names are linked to treatments from the Manual


Key to HELIANTHUS

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Annual; petioles ± long, well developed
 
    2. Phyllaries generally > 4 mm wide, abruptly acuminate; leaf base cordate
H. annuus
    2' Phyllaries generally < 4 mm wide, acute or gradually acuminate; leaf base generally truncate to wedge-shaped
 
      3. Central chaff scales tipped with stiff white hairs
 
        4. Herbage and phyllaries densely gray-canescent
H. niveus subsp. canescens
        4' Herbage and phyllaries strigose, green
H. petiolaris subsp. petiolaris
      3' Central chaff scales glabrous or very finely appressed-hairy
 
        5. Chaff scales awn-tipped, >> disk flowers, glabrous; leaves green
H. bolanderi
        5' Chaff scales acute, = or slightly > disk flowers, rough-hairy; leaves densely white-hairy
2 H. niveus subsp. tephrodes
  1' Perennial; leaves sessile (or petiole < 3 cm)
 
    6. Disk corolla lobes red to purple
 
      7. Leaves blue-green, ± glabrous, margins often crinkled; roots horizontal, rhizome-like; ligules generally < or = 1 cm; phyllaries < disk
H. ciliaris
      7' Leaves green or densely white-hairy, margins generally flat; plant taprooted; ligules generally >> 1 cm; phyllaries = or > disk
 
        8. Leaves green, short-rough-hairy; phyllaries not > disk
2 H. gracilentus
        8' Leaves densely white-hairy; phyllaries = or slightly > disk
2 H. niveus subsp. tephrodes
    6' Disk corolla lobes yellow
 
      9. Phyllaries not > disk, obtuse to acute
2 H. gracilentus
      9' Phyllaries > disk, acuminate
 
        10. Phyllaries generally 12–18; most leaves opposite, generally < 12 cm; plants tap-rooted
H. cusickii
        10' Phyllaries generally > 25; most leaves alternate, often > 15 cm: plants rhizomed
 
          11. Leaf blades folded along midvein; inflorescence generally raceme-like
H. maximilianii
          11' Leaf blades ± flat; inflorescence flat-topped or rounded
 
            12. Phyllaries generally 3+ mm wide, folded back at maturity; stem generally glabrous
H. californicus
            12' Phyllaries generally < 3 mm wide, ± erect at maturity; stem generally ± hairy
H. nuttallii
              13. Phyllaries sparsely hairy; leaf blade short-rough-hairy below
subsp. nuttallii
              13' Phyllaries densely hairy; leaf blade finely tomentose below
subsp. parishii


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