The Jepson Manual: Higher Plants of California

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Names are linked to treatments from the Manual


Key to GILIA Key to Groups

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Cobwebby (fine, kinky) hairs in leaf axils or elsewhere; stem ± scapose; basal leaves generally in rosette
Group 1
  1' Cobwebby hairs 0; stem ± scapose or not; basal rosette present or 0
Group 2

Key to GILIA Group 1: Cobwebby hairs present; st +- scapose; basal rosette gen present (sect. Arachnion)

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Calyx, pedicel densely glandular (rarely cobwebby); flowers, fruits generally in clusters; calyx ribs wider than membrane
G. brecciarum
    2. Stamens long-exserted; corolla 10–20 mm, upper throat white with yellow spots
subsp. neglecta
    2' Stamens slightly exserted; corolla 5–11 mm, upper throat ± purple
 
      3. Branches spreading to nearly erect; corolla throat yellow with purple veins
subsp. brecciarum
      3' Branches long-decumbent; corolla throat generally all purple
subsp. jacens
  1' Calyx, pedicel barely or not glandular; flowers, fruits generally not in clusters; calyx ribs narrower than membrane
 
    4. Cauline leaves clasping or expanded at base
 
      5. Corolla 6–12 mm, 1–2 × calyx
 
        6. Stem green (not glaucous), slightly cobwebby below middle; upper leaves generally deeply toothed
G. modocensis
        6' Stem glaucous, glabrous below middle; upper leaves generally not deeply toothed
 
          7. Stamens, style exserted to middle of corolla lobes; pollen blue; generally s of Transverse Ranges
G. diegensis
          7' Stamens, style exserted to base of corolla lobes; pollen white; generally n of Transverse Ranges
G. sinuata
      5' Corolla 10–35 mm, 2–7 × calyx
G. latiflora
        8. Stem cobwebby below middle
 
          9. Corolla tube stout, throat > 7 mm wide, corolla generally 3 × lobes-> subsp. 2davyi
          9' Corolla tube slender, throat < 7 mm wide, corolla 4–6 × lobes
subsp. elongata
        8' Stem glabrous and glaucous below middle
 
          10. Flowers in loose clusters; corolla purple to mid-throat or higher-> subsp. 2davyi
          10' Flowers not in clusters; corolla purple to base of throat
 
            11. Calyx 2–4 mm; corolla 9–16 mm
subsp. cuyamensis
            11' Calyx 4–5 mm; corolla 15–22 mm
subsp. latiflora
    4' Cauline leaves not clasping or expanded at base
 
      12. Calyx glabrous or cobwebby (not glandular)
 
        13. Stamens = or > corolla lobes; calyx membrane generally purple or purple-spotted
 
          14. Basal leaves densely cobwebby; calyx membrane generally dull purple-spotted; High Sierra Nevada, w Great Basin Floristic Province
G. salticola
          14' Basal leaves glabrous or slightly cobwebby; calyx membrane bright purple-spotted; Mojave Desert
G. aliquanta
            15. Corolla lobes = tube + throat
subsp. aliquanta
            15' Corolla lobes generally 1/2 tube + throat
subsp. breviloba
        13' Stamens < corolla lobes; calyx membrane generally not colored
 
          16. Basal leaf lobes spreading; corolla throat yellow or yellow-spotted
 
            17. Corolla 4–7 mm; throat yellow-spotted, < or = lobes; calyx lobes short-pointed
G. clokeyi
            17' Corolla 7–12 mm; throat yellow, = or > lobes; calyx lobes acuminate
G. ophthalmoides
          16' Basal leaf lobes generally ascending; corolla throat blue in upper 1/2, yellow below
G. ochroleuca
            18. Stem glabrous and glaucous below inflorescence
 
              19. Corolla 8–14 mm; style > stamens
subsp. bizonata
              19' Corolla 4–6 mm; style < or = stamens
subsp. ochroleuca
            18' Stem generally cobwebby or glandular below inflorescence (if glabrous, not glaucous)
 
              20. Plant tall; basal leaves entire to generally 1-pinnate; corolla pale; s San Bernardino Mountains, Peninsular Ranges
subsp. exilis
              20' Plant low, spreading; basal leaves 2-pinnate; corolla bright; San Gabriel Mountains, Tehachapi Mountain Area
subsp. vivida
      12' Calyx glandular (rarely cobwebby in young plants)
 
        21. Lobes of basal leaves toothed on both sides
 
          22. Stamens < corolla lobes; corolla throat bluish in upper 1/2, yellow in lower 1/2
G. cana
            23. Adjacent pedicels widely spreading
 
              24. Corolla throat conic, tube 2–4 × calyx
subsp. speciformis
              24' Corolla throat cup-shaped, tube generally < 2 × calyx
subsp. triceps
            23' Adjacent pedicels narrowly spreading
 
              25. Fruit narrowly ovoid, generally < 5 mm; basal leaves cobwebby-matted
subsp. cana
              25' Fruit widely ovoid, generally > 5 mm; basal leaves cobwebby, not matted
 
                26. Corolla tube = throat; stamens unequal, longest well exserted
subsp. bernardina
                26' Corolla tube 3–6 × throat; stamens equal, slightly exserted
subsp. speciosa
          22' Stamens generally > corolla lobes; corolla throat yellow or white
G. leptantha
            27. Corolla tube 2–4 × calyx
 
              28. Corolla lobes 1–3 mm wide; San Bernardino Mountains
subsp. leptantha
              28' Corolla lobes 3–5 mm wide; s Sierra Nevada
subsp. purpusii
            27' Corolla tube 1–2 × calyx
 
              29. Corolla throat conic; stamens > corolla lobes
subsp. pinetorum
              29' Corolla throat cup-shaped; stamens < corolla lobes
subsp. transversa
        21' Lobes of basal leaves linear, entire or toothed on 1 side
 
          30. Inflorescence showy; corolla 10–22 mm, unspotted below lobes
G. tenuiflora
            31. Corolla tube stout, lobes 4–7 mm wide
 
              32. Corolla throat > lobes; mainland
subsp. amplifaucalis
              32' Corolla throat < lobes; Channel Islands
subsp. hoffmannii
            31' Corolla tube slender, lobes 2–4 mm wide
 
              33. Branches decumbent; calyx densely glandular; stamens exserted to base of lobes
subsp. arenaria
              33' Branches spreading, calyx slightly glandular; longest stamens exserted beyond middle of lobes
subsp. tenuiflora
          30' Inflorescence not showy; corolla 3–12 mm, generally spotted below lobes (G. inconspicua complex)
 
            34. Longest stamens exserted > mid-lobe
 
              35. Leaf lobes ascending; corolla 3–4 × lobes; Great Basin Floristic Province
G. inconspicua
              35' Leaf lobes spreading; corolla 2–3 × lobes; s Sierra Nevada Foothills, Tehachapi Mountain Area
G. interior
            34' Longest stamens exserted < mid-lobe
 
              36. Fruit 2 × calyx, valves not detaching; leaf lobes < 1 mm wide, ascending
G. minor
              36' Fruit = calyx, valves detaching; leaf lobes 1–2 mm wide, spreading
 
                37. Branches decumbent; corolla without spots, lobe tips obtuse
G. malior
                37' Branches erect to spreading; corolla with purple spots below lobes, lobe tips acute
 
                  38. Lobes of basal leaves 2–3 mm; fruit 3–4 mm; South Coast Ranges
G. austro-occidentalis
                  38' Lobes of basal leaves 3–11 mm; fruit 5–6 mm; Mojave Desert
G. transmontana

Key to GILIA Group 2: Cobwebby hairs 0; st +- scapose or not; basal rosette sometimes 0

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Sepals generally fused below middle or at base; pollen yellow (sect. Giliastrum)
 
    2. Principal leaves sharp-toothed, holly-like; plant > 10 cm; corolla pink
 
      3. Annual; leaf teeth acuminate
G. latifolia
      3' Perennial; leaf teeth needle-like
G. ripleyi
    2' Principal leaves not sharp-toothed, not holly-like; plant < 9 cm; corolla yellow or white
 
      4. Leaf linear; corolla yellow
G. filiformis
      4' Leaf oblong, lanceolate, or narrowly oblanceolate; corolla mostly white
 
        5. Plant < 3 cm, stem not glandular; leaf 2–5 mm; calyx membrane ciliate; Sonoran Desert
G. maculata
        5' Plant > 3 cm, stem glandular; leaf 6–20 mm; calyx membrane not ciliate; Great Basin Floristic Province, Mojave Desert
 
          6. Corolla throat spotted, tube exserted from calyx
G. campanulata
          6' Corolla throat not spotted, tube included in calyx
 
            7. Lower leaves white-hairy; corolla 4–7 mm; seeds 10–15 per chamber
G. inyoensis
            7' Lower leaves generally glabrous; corolla < 3 mm; seeds 1 per chamber
G. tenerrima
  1' Sepals generally fused to middle or higher; pollen blue, yellow, or white
 
    8. Plant glandular-puberulent; upper leaves generally entire
 
      9. Leaves generally cauline; inflorescence not scapose; pollen blue; seeds gelatinous when wet (sect. Kelloggia)
 
        10. Corolla 6–8 mm, lobes bluish white; branches, leaves ascending
G. capillaris
        10' Corolla 5–20 mm, lobes pink; branches, leaves spreading
 
          11. Calyx glandular; corolla tube, throat yellow with reddish streaks
G. sinistra
            12. Corolla 9–20 mm; style exserted
subsp. pinnatisecta
            12' Corolla 5–9 mm; style included
subsp. sinistra
          11' Calyx glabrous; corolla tube yellow, throat purple or yellow
G. leptalea
            13. Corolla 8–15 mm, throat 2–5 mm, yellow, sometimes with short purple marks
subsp. bicolor
            13' Corolla 13–21 mm, throat 6–8 mm, purple
subsp. leptalea
      9' Leaves in basal rosette; inflorescence generally ± scapose; pollen yellow or white; seeds not gelatinous when wet (sect. Giliandra)
 
        14. Lower leaves generally 2-pinnately lobed; corolla tube glandular-puberulent, lobes wavy-margined
G. hutchinsifolia
        14' Lower leaves generally 1-pinnately lobed or toothed; corolla tube glabrous, lobes not wavy-margined
 
          15. Corolla tube thread-like, exserted from calyx
 
            16. Corolla lobes truncate and short-pointed; throat wider than tube
G. leptomeria
            16' Corolla lobes 3-toothed; throat narrower than tube
G. triodon
          15' Corolla tube stout, included or slightly exserted from calyx
 
            17. Corolla 2–4 mm, throat generally included; pedicels in fruit spreading or recurved; fruit spheric, generally < calyx
G. micromeria
            17' Corolla 4–7 mm, throat exserted; pedicels in fruit nearly erect; fruit ovoid, generally > calyx
 
              18. Leaf axis strap-shaped; basal leaves generally without sand grains on upper surface
G. lottiae
              18' Leaf axis linear; basal leaves dotted with sand grains on both surfaces
G. subacaulis
    8' Plant hairy on leaf or stem; upper leaves lobed or toothed
 
      19. Leaf lobes generally not linear; flowers not in heads or clusters; pedicel glands large, generally flat-topped, subsessile (sect. Saltugilia)
 
        20. Calyx gland-dotted
 
          21. Hairs on basal leaves translucent, straight or curved, gland-tipped; spots 0 in corolla
G. scopulorum
          21' Hairs on basal leaves white, sharply bent, non-glandular; purple spots below each corolla lobe
G. stellata
        20' Calyx generally glabrous
 
          22. Corolla 5–13 mm, lavender or white
 
            23. Stamens < corolla lobes; corolla lobes 1–1.5 × throat, throat not purple-marked
G. australis
            23' Stamens generally > corolla lobes; corolla lobes 2–3 × throat, throat purple-marked below each lobe
G. caruifolia
          22' Corolla 10–36 mm, generally pink
G. splendens
            24. Corolla tube 7–18 mm, 2–5 × calyx
subsp. grantii
            24' Corolla tube 4–10 mm, 1–2 × calyx
subsp. splendens
      19' Leaf lobes ± linear; flowers generally in heads or clusters; pedicel glands, if present, minute, spheric, stalked (sect. Gilia)
 
        25. Inflorescence axis generally glandular; corolla throat yellow, sometimes purple-spotted
 
          26. Corolla throat not spotted
 
            27. Corolla 6–9 mm; throat cup-shaped; leaves 1–2-pinnate
G. angelensis
            27' Corolla 8–14 mm, throat conic; leaves 2–3-pinnate
G. nevinii
          26' Corolla throat purple-spotted
 
            28. Corolla 8–19 mm, throat well exserted from calyx
G. tricolor
              29. Flowers in loose clusters or solitary; corolla 7–13 mm
subsp. diffusa
              29' Flowers generally in dense clusters; corolla 10–19 mm
subsp. tricolor
            28' Corolla 7–11 mm, throat partly included in calyx
 
              30. Calyx in fruit 5–8 mm, generally < or = fruit
G. clivorum
              30' Calyx in fruit 8–13 mm, generally >> fruit
G. millefoliata
        25' Inflorescence axis generally glabrous or tomentose; corolla throat neither yellow nor spotted
 
          31. Flowers generally in hemispheric heads or clusters; corolla throat generally > tube; stamens < or = corolla lobes
G. achilleifolia
            32. Corolla 10–20 mm; heads 8–25-flowered; pedicels 1–2 mm
subsp. achilleifolia
            32' Corolla 5–10 mm; clusters 2–7-flowered; pedicels 1–30 mm
subsp. multicaulis
          31' Flowers in spheric heads; corolla throat < or = tube; stamens = or > corolla lobes
G. capitata
            33. Corolla 5–8 mm, lobes linear or narrowly oblong; calyx lobes acute, erect
 
              34. Heads glabrous or sparsely hairy to glandular at base
 
                35. Calyx membrane colorless; corolla lobes < 1 mm wide; seeds generally 1–6
subsp. capitata
                35' Calyx membrane blue-violet; corolla lobes 1–2 mm wide; seeds 10–25
subsp. pacifica
              34' Heads tomentose at base
 
                36. Heads 12–25 mm wide; corolla pale blue-violet; Sierra Nevada
subsp. mediomontana
                36' Heads 20–35 mm wide; corolla bright blue-violet; North Coast
subsp. tomentosa
            33' Corolla 7–13 mm, lobes oblong; calyx lobes acuminate, tips recurved
 
              37. Heads sparsely hairy to glabrous at base; flowers short-pedicelled; s California
subsp. abrotanifolia
              37' Heads tomentose at base; flowers generally sessile; especially n&c California
 
                38. Corolla bright blue-violet; plant with skunk-like odor — n Central Coast
subsp. chamissonis
                38' Corolla pale blue-violet; plant not with skunk-like odor
 
                  39. Inflorescence 10–20 mm wide; corolla lobes ± 2 mm wide; Sierra Nevada Foothills
subsp. pedemontana
                  39' Inflorescence 20–30 mm wide; corolla lobes ± 3 mm wide; Great Central Valley, Inner South Coast Ranges
subsp. staminea


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