The Jepson Manual: Higher Plants of California

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Names are linked to treatments from the Manual


Key to CAMISSONIA

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Ovary tip (below hypanthium, above seeds) slender, sterile, 5–180 mm; stem ± 0 (sect. Tetrapteron)
 
    2. Annual; fruit winged
 
      3. Leaf linear to narrowly lanceolate or oblanceolate; petals 5–18 mm; hypanthium 1.6–3.2 mm; plant densely spreading-hairy
C. graciliflora
      3' Leaf narrowly oblanceolate; petals 2–3.5 mm; hypanthium 0.8–1.3 mm; plant strigose and sparsely spreading-hairy
C. palmeri
    2' Perennial; fruit not winged
 
      4. Leaf pinnately lobed; plant hairs ± dense, spreading or appressed; hypanthium 4–8.5 mm; stigma > anthers
C. tanacetifolia
        5. Plant hairs spreading, dense; pollen > 10% 4-angled
subsp. quadriperforata
        5' Plant hairs spreading or appressed; pollen < 5% 4-angled
subsp. tanacetifolia
      4' Leaf not pinnately lobed (or plant ± glabrous); hypanthium 1.5–3 mm; stigma ± = or slightly > anthers
 
        6. Leaf hairs short, erect; fruit subcylindric (swollen by seeds), papery
C. ovata
        6' Leaf subglabrous; fruit 4-angled, barely swollen by seeds, leathery
C. subacaulis
  1' Ovary without a slender, sterile tip; stem present (or plant immature)
 
    7. Seed wing thick, with club-shaped hairs; petals white, bases yellow (sect. Chlismiella)
C. pterosperma
    7' Seed wing 0; petals white, cream, yellow, lavender, rarely red
 
      8. Fruit pedicelled, not coiled, twisted, or wavy; seeds in 2 rows per chamber
 
        9. Hypanthium 4.5–40 mm; plant with cauline, simple leaves (rosette 0 or poorly developed) (sect. Lignothera)
 
          10. Hypanthium 18–40 mm; inflorescence open; sepals 8–15 mm
C. arenaria
          10' Hypanthium 4.5–14 mm; inflorescence dense; sepals 3–9 mm
C. cardiophylla
            11. Hairs generally spreading (sometimes some glandular); hypanthium 4.5–12 mm
subsp. cardiophylla
            11' Hairs generally glandular (some spreading, nonglandular); hypanthium 9–14 mm
subsp. robusta
        9' Hypanthium 0.4–8 mm; plant generally with well developed basal rosette (sometimes also cauline leaves); leaves generally 1-pinnate (sect. Chylismia)
 
          12. Petals lavender, bases generally yellow, lavender-dotted; rosette poorly developed
C. heterochroma
          12' Petals yellow or white (sometimes fading purple or red or bases purple- or red-dotted); rosette well developed
 
            13. Fruit generally < 2 mm wide, ± same width throughout
 
              14. Sepals 1.5–4 mm; hypanthium 1–1.5 mm; stigma ± = anthers; inflorescence erect, buds drooping; leaflets < 30 mm
C. walkeri subsp. tortilis
              14' Sepals 5–9 mm; hypanthium 3–8 mm; stigma > anthers; inflorescence nodding; leaflets generally < 10 mm or 0
C. brevipes
                15. Flower bud reflexed; petals generally fading red
subsp. arizonica
                15' Flower bud not reflexed; petals not fading red
 
                  16. Plant hairs spreading; sepals in bud with free subterminal tips; petals not red-dotted
subsp. brevipes
                  16' Plant strigose (rarely also with spreading hairs below); sepals in bud without free tips; petals generally red-dotted
subsp. pallidula
            13' Fruit > 2 mm wide, wider toward tip
 
              17. Pedicel and fruit becoming reflexed; petals yellow
C. munzii
              17' Pedicel and fruit spreading or ascending; petals white or yellow
C. claviformis
                18. Plant spreading-hairy below; free tips of sepals in bud conspicuous; petals yellow
subsp. peirsonii
                18' Plant strigose or glabrous below; free tips of sepals in bud 0 or inconspicuous (except subsp. funerea); petals white or yellow
 
                  19. Petals yellow
 
                    20. Plant strigose above, also glandular or not; free tips of sepals in bud 0 or short
subsp. yumae
                    20' Plant glabrous above or glandular-hairy; free tips of sepals in bud 0
 
                      21. Terminal leaflet ovate to subcordate, lateral leaflets few–many; flower generally opening at dawn
subsp. cruciformis
                      21' Terminal leaflet lanceolate, lateral leaflets 0–few; flower opening at dusk
subsp. lancifolia
                  19' Petals white (rarely pale yellow in subsp. claviformis)
 
                    22. Lateral leaflets generally well developed
 
                      23. Plant strigose above, rarely glandular-hairy
subsp. aurantiaca
                      23' Plant ± glabrous above (or with some glandular hairs)
subsp. claviformis
                    22' Lateral leaflets generally 0 or poorly developed
 
                      24. Free tips of sepals in bud conspicuous; plant generally strigose above
subsp. funerea
                      24' Free tips of sepals in bud 0 or minute; plant ± glabrous above or generally with some glandular hairs
subsp. integrior
      8' Fruit ± sessile (except some C. kernensis), often coiled, twisted and wavy; seeds in 1 row per chamber
 
        25. Petals white; flower opening at dusk (sect. Eremothera)
 
          26. Fruit of ± same width throughout; flowers at lower nodes 0
 
            27. Sepals 1.5–2.5 mm; petals 1.8–3 mm; stigma ± surrounded by anthers
C. chamaenerioides
            27' Sepals 4–6 mm; petals 3.5–7 mm; stigma exceeding anthers
C. refracta
          26' Fruit base wider than tip; flowers at lower nodes present or 0
 
            28. Inflorescence erect; petals 0.8–1.3 mm; sepals 0.8–1.8 mm
C. minor
            28' Inflorescence nodding; petals 3–7.5 mm; sepals (2.7)4–8 mm
C. boothii
              29. Rosette present at time of first flower (spring); bracts not leaf-like, ± inconspicuous; plant ± glabrous (or hairs not spreading)
 
                30. Fruit 2–3.8 mm wide at base, woody, curved outward but not downward
subsp. condensata
                30' Fruit 1–2.3 mm wide at base, not woody, curved outward or downward
 
                  31. Fruit 1.7–2.3 mm wide at base, curved outward
subsp. decorticans
                  31' Fruit 1–1.6 mm wide at base, generally curved downward
subsp. desertorum
              29' Rosette generally withered by time of first flower (late spring, summer); bracts leaf-like, conspicuous; plant hairs often spreading
 
                32. Plant strigose, also with rarely spreading or glandular hairs; seeds all minutely pitted
subsp. alyssoides
                32' Plant hairs spreading, some glandular; seeds of 2 kinds, minutely pitted and coarsely papillate
 
                  33. Plant generally 15–40 cm; cauline leaf lanceolate to narrowly ovate, serrate
subsp. boothii
                  33' Plant 5–20 cm; cauline leaf narrowly lanceolate to lanceolate, ± entire to minutely serrate
subsp. intermedia
        25' Petals yellow; flower opening near dawn
 
          34. Plant straight, slender, erect; leaf sharply pinnately lobed; petal bases ± red-flecked; fruit reflexed (sect. Eulobus)
C. californica
          34' Plant not simultaneously straight, slender, and erect; leaf entire to toothed; petal bases with 0–2 red dots; fruit not reflexed
 
            35. Leaves closely spaced at stem tips; fruit very flat, 5–10 mm (sect. Nematocaulis)
C. andina
            35' Leaves well spaced throughout; fruit 4-angled or cylindric, often swollen by seeds, generally > 10 mm
 
              36. Fruit cylindric, ± swollen by seeds; flowers 0 at lower nodes; leaf generally linear to narrowly elliptic; seeds shiny, often < 1 mm (sect. Camissonia)
 
                37. Sepals all separating when flower opens
 
                  38. Petals 8–18 mm; sepals 5–11 mm; stigma exceeding anthers
C. kernensis
                    39. Plant somewhat open, sparsely spreading-hairy, sparsely glandular-hairy leaves not clustered at base; pedical 0–5 mm
subsp. gilmanii
                    39' Plant compact, densely spreading-hairy, sparsely glandular-hairy; leaves clustered at base; pedicel 3–15 mm
subsp. kernensis
                  38' Petals 1.8–4 mm; sepals < 3.8 mm; stigma ± surrounding anthers
 
                    40. Rosette 0; leaf subentire; plant ± glabrous or minutely strigose, generally sparsely glandular-hairy, rarely sparsely spreading-nonglandular-hairy
C. parvula
                    40' Rosette generally ± 0; leaf serrate; plant hairs generally spreading, some glandular
 
                      41. Hypanthium 1.3–3 mm; sepals 2.2–3.8 mm; fruit (18)26–50 mm
C. pubens
                      41' Hypanthium 0.8–1.6 mm; sepals 1.2–2 mm; fruit 18–32 mm
C. pusilla
                37' Sepals generally remaining adherent in pairs when flower opens
 
                  42. Stigma exceeding anthers; sepals 3–8(12) mm; petals (3.5)5–15 mm
 
                    43. Leaf lanceolate to narrowly ovate or elliptic, subentire; sepals < 4.2 mm
2 C. sierrae subsp. sierrae
                    43' Leaf linear to narrow-elliptic or -oblanceolate, minutely or coarsely serrate; sepals 3–8(12) mm
C. campestris
                      44. Stem generally erect; leaf minutely serrate
subsp. campestris
                      44' Stem generally decumbent; leaf coarsely serrate
subsp. obispoensis
                  42' Stigma generally surrounded by anthers; sepals 1.6–4(5.5) mm; petals 2–7 mm
 
                    45. Leaf entire (rarely with 1–2 small teeth); inflorescence densely minutely strigose
C. integrifolia
                    45' Leaf minutely serrate (C. sierrae sometimes with only several small teeth); inflorescence hairs glandular to spreading, sometimes minutely strigose and glandular
 
                      46. Leaf lanceolate to narrowly ovate or elliptic, base obtuse or rounded
C. sierrae
                        47. Sepals 1.2–3 mm; petal bases not red-dotted; style 2.8–5 mm
subsp. alticola
                        47' Sepals 3–4.2 mm; petal bases with 2 red dots; style 4.5–7 mm
subsp. sierrae
                      46' Leaf linear to narrowly elliptic, base acute or long-tapered
 
                        48. Plant minutely strigose (or some hairs spreading and glandular, or sometimes coarsely spreading and nonglandular toward base); < 10% of pollen grains 4-angled
C. strigulosa
                        48' Plant hairs spreading, fine (in inflorescence often glandular); > 30% of pollen grains 4-angled
 
                          49. Sepals (3)3.8–5.5 mm; petals (4)4.5–7 mm
C. lacustris
                          49' Sepals 1.6–4 mm; petals 2–4(5) mm
 
                            50. Leaf generally bluish green; hypanthium 1.2–2.3 mm; plant hairs coarse; generally > 30% of pollen grains 4-angled; widespread
C. contorta
                            50' Leaf not (or slightly) bluish green; hypanthium ± 1.2 mm; plant hairs fine; < 10% of pollen grains 4-angled; Inner South Coast Ranges
C. benitensis
              36' Fruit 4-angled (at least when dry, except C. hirtella), not swollen by seeds; flowers generally at lower (through upper) nodes; leaf generally lanceolate to narrowly elliptic or narrowly ovate; seeds dull, often > 1 mm (sect. Holostigma)
 
                51. Perennial or subshrub; ± coastal
C. cheiranthifolia
                  52. Perennial; hairs rarely silvery; style 6–9 mm; petals 6–11 mm; stigma generally ± surrounding anthers
subsp. cheiranthifolia
                  52' Subshrub; hairs silvery; style 13–23 mm; petals (10)12–20 mm; stigma exceeding anthers
subsp. suffruticosa
                51' Annual or short-lived perennial herb; inland or coastal
 
                  53. Stigma exceeding anthers
C. bistorta
                  53' Stigma generally surrounded by anthers
 
                    54. Fruit stout (2.8–3.5 mm wide at base), deeply grooved
C. guadalupensis subsp. clementina
                    54' Fruit not especially stout (0.7–2.2 mm wide at base), not deeply grooved
 
                      55. > 25% of pollen grains 4–5-angled; very localized species; 2n=42
 
                        56. Inflorescence hairs nonglandular; 25–60% of pollen grains 4–5-angled — Outer South Coast Ranges (Monterey Co.)
C. luciae
                        56' Inflorescence hairs both glandular and other types; 70–100% of pollen grains 4–5-angled
 
                          57. Fruit 1.3–1.6 mm wide, subcylindric (when dry ± 4-angled); Outer South Coast Ranges (Monterey, San Luis Obispo cos.)
C. hardhamiae
                          57' Fruit 1.5–2 mm wide, 4-angled; Channel Islands, Peninsular Ranges
C. robusta
                      55' < 5% of pollen grains 4–5-angled; localized or widespread species; 2n=14,28
 
                        58. Fruit conspicuously 4-angled, 1.8–2.2 mm wide — c&s South Coast
C. lewisii
                        58' Fruit cylindric or weakly 4-angled when fresh, 0.7–1.8 mm wide
 
                          59. Upper petioles < 25 mm; fruit < 5-coiled
C. ignota
                          59' Upper petioles 0 or < 2 mm; fruit straight to 3-coiled
 
                            60. Plant strigose, grayish — s San Joaquin Valley, Desert
C. pallida
                              61. Petals 6.5–13 mm; hypanthium 3.8–4.2 mm; style 6.5–10.5 mm
subsp. hallii
                              61' Petals 2–6.5 mm; hypanthium 1–3 mm; style 2.1–6.5 mm
subsp. pallida
                            60' Plant not strigose, sometimes grayish
 
                              62. Fruit 0.7–0.9 mm wide; upper leaves narrowly ovate to ovate; stem ascending
C. hirtella
                              62' Fruit 0.8–1.8 mm wide; upper leaves narrowly lanceolate to narrowly ovate; stem ± erect or decumbent
 
                                63. Stem generally decumbent; upper leaves generally narrowly lanceolate; inflorescence hairs generally nonglandular
C. micrantha
                                63' Stem ± erect; upper leaves lanceolate to narrowly ovate; inflorescence generally with glandular hairs
 
                                  64. Plant grayish with spreading hairs; hypanthium 2–3.8 mm; sepals 3.2–8 mm
C. confusa
                                  64' Plant with spreading hairs, but not conspicuously grayish; hypanthium 1.2–2 mm; sepals 1–2.5 mm
C. intermedia


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