The Jepson Manual: Higher Plants of California

    THIS PAGE IS NO LONGER UPDATED
    AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY
  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

Names are linked to treatments from the Manual


Key to BRASSICACEAE Key to Groups

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Fruit generally at least 4 × longer than wide, generally linear
Group 1
  1' Fruit < 4 × longer than wide, not linear
 
    2. Fruit round to ± flat in X-section and sometimes inflated
Group 2
    2' Fruit flat, not inflated
 
      3. Fruit 1-seeded, 1-chambered, indehiscent
Group 3
      3' Fruit generally 2±seeded, 2-chambered, generally dehiscent
 
        4. Fruit flat perpendicular to septum
Group 4
        4' Fruit flat parallel to septum
Group 5

Key to BRASSICACEAE Group 1

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Leaf hairs branched (sometimes also simple)
 
    2. Leaf deeply lobed or compound; fruit < 3 cm
 
      3. Leaf segments rigid, often sharp-pointed; fruit often flat perpendicular to septum
POLYCTENIUM (no key)2
      3' Leaf segments ± flexible, not sharp-pointed; fruit ± cylindric or ± flat parallel to septum
 
        4. Fruit ± cylindric, valves midribbed ± to tip; pedicel generally thread-like; petals yellow or whitish
DESCURAINIA
        4' Fruit generally flat, valves midribbed only at base; pedicel linear, width nearly = fruit width; petals white to purplish
SIBARA
    2' Leaf entire to generally shallowly lobed; fruit often > 3 cm
 
      5. Fruit flat
 
        6. Hairs appressed, forked to multibranched; petals generally > 15 mm, blade much wider than claw
ERYSIMUM
        6' Hairs appressed and not, simple to multibranched; petals < 15 mm, blade little wider than claw
 
          7. Seeds 1 row per chamber
 
            8. Fruit not long-tapered; taproot often ± slender; basal leaves entire or dentate, generally < 10 cm; seed often winged
ARABIS
            8' Fruit long-tapered to tip; taproot very thick; basal leaves entire, 3–15 cm; seed unwinged
PHOENICAULIS (no key)
          7' Seeds 2 rows per chamber
 
            9. Fruit 5–7 mm wide, tapered, sharp-pointed; seed coat silvery
ANELSONIA (no key)2
            9' Fruit < 5 mm wide (or tip not tapered and sharp-pointed); seed coat not silvery
 
              10. Fruit generally > 3 cm, > 10 × longer than wide; petals not yellow
ARABIS
              10' Fruit generally < 2 cm, < 10 × longer than wide; petals often yellow
DRABA
      5' Fruit ± flat to ± 4-sided or round in X-section
 
        11. Stigma lobes pointed, decurrent
MALCOLMIA (no key)
        11' Stigma lobes not pointed, decurrent or not
 
          12. Flower showy (petals generally > 15 mm, blade much wider than claw); fruit generally > 4 cm (inflated if shorter)
 
            13. Leaf and stem hairs appressed, forked to multibranched; petals often yellow or orange
ERYSIMUM
            13' Leaf and stem hairs appressed and not, simple to multibranched; petals not yellow or orange
 
              14. Fruit < 3 cm, inflated; style 4–8 mm; stigma lobes not decurrent
AUBRIETA (no key)
              14' Fruit > 3 cm, not inflated; style < 4 mm; stigma lobes ± decurrent
 
                15. Leaf dentate; leaf hairs ± sparse, simple and forked, fruit hairs generally 0
HESPERIS (no key)
                15' Leaf ± entire; leaf and fruit hairs dense, multibranched, rarely also simple
MATTHIOLA (no key)
          12' Flower not showy (petals generally < 10 mm, blade little wider than claw); fruit generally < 4 cm (except Arabis glabra), not inflated
 
            16. Hairs appressed, forked to multibranched; petals yellow
ERYSIMUM
            16' Hairs appressed or not, simple to multibranched; petals white or cream-colored, rarely lilac
 
              17. Fruit 4–10 cm; upper stem leaves clasping
Arabis glabra
              17' Fruit < 4 cm; upper stem leaves not clasping
 
                18. Annual; fruit glabrous; seeds ± 1 row per chamber
ARABIDOPSIS (no key)
                18' Biennial or perennial herb; fruit sometimes hairy; seeds 2 rows per chamber
HALIMOLOBOS
  1' Leaf hairs 0 or simple (rarely branched in Caulanthus)
 
    19. Fruit stalk above pedicel 1–3 cm; stamens ± equal
STANLEYA
    19' Fruit stalk above pedicel 0(–1) cm; stamens generally 4 long, 2 short
 
      20. Calyx ± urn- or flask-shaped, nearly closed at full flower, often brightly colored; petals generally strap-shaped (except Streptanthella)
 
        21. Fruit generally ± cylindric or ± flat parallel to septum; seed generally unwinged
 
          22. Sepals 3–18 mm, generally not green; petals 5–23 mm, blade base as wide as or narrower than claw; stigma lobes generally 2; fruit 1.5–18 cm, ± cylindric or ± flat; seed 1–4 mm, generally ± brown, wing sometimes narrow at 1 end or 0
CAULANTHUS
          22' Sepals 1–11 mm, greenish or not; petals 3–15 mm, blade base sometimes much wider than claw; stigma lobes 0 or 2; fruit 1–9 cm, ± cylindric; seed < 2 mm, brownish or yellowish, wing 0
GUILLENIA
        21' Fruit generally flat parallel to septum; seed generally winged
 
          23. Fruit pendent, valves persistent to beak
STREPTANTHELLA (no key)
          23' Fruit ascending or pendent, valves dehiscent, falling, beak 0
STREPTANTHUS
      20' Calyx not urn- or flask-shaped, fully opening, generally greenish; petals generally clawed or spoon-shaped, sometimes linear to obovate
 
        24. Bracts subtending at least several flowers per inflorescence
 
          25. Fruit ± 4-sided
ERUCA (no key)STRUM*
          25' Fruit flat
TROPIDOCARPUM
        24' Bracts generally 0 (may subtend a few lower flowers or main branches)
 
          26. Fruit not opening by valves
 
            27. Fruit flat, winged, oblong to oblanceolate, indehiscent, not breaking into segments
ISATIS (no key)2
            27' Fruit ± flat or ± round in X-section, unwinged, not oblong to oblanceolate, generally breaking into transverse segments
 
              28. Fruit beak narrow, > 5 mm; fruit breaking into 2–several 0–5-seeded segments
 
                29. Plant glandular; fruit often upcurved, not grooved
CHORISPORA (no key)
                29' Plant not glandular; fruit straight, grooved below
RAPHANUS
              28' Fruit beak either wide or < 5 mm; fruit breaking into 2 unlike segments, each generally 1-seeded
 
                30. Plant generally glabrous; leaf fleshy; ocean dunes; petals white to lavender; fruit 1–3 cm
CAKILE
                30' Plant coarsely stiff-hairy; leaf not fleshy; disturbed areas (not dunes); petals yellow; fruit < 1 cm
RAPISTRUM (no key)
          26' Fruit opening by valves
 
            31. Fruit beak prominent, generally nearly as wide as valves
 
              32. Seeds 2 rows per chamber; petals generally prominently dark-veined
ERUCA
              32' Seeds 1 row per chamber; petals not dark-veined
 
                33. Fruit valves with 1 midvein
BRASSICA
                33' Fruit valves with 3–7 veins, especially when young
 
                  34. Fruit appressed to stem, beak base abruptly swollen
HIRSCHFELDIA (no key)
                  34' Fruit not appressed to stem, beak base generally not abruptly swollen
SINAPIS
            31' Fruit beak generally 0 or obscure (except Brassica), narrower than valves
 
              35. Fruit generally ± flat
 
                36. Fruit clearly stalked above receptacle
THELYPODIUM
                36' Fruit ± not stalked above receptacle
 
                  37. Fruit valves without prominent midrib
 
                    38. Fruit valves generally opening elastically, sometimes coiling from base
CARDAMINE
                    38' Fruit valves not opening elastically
SIBARA
                  37' Fruit valves with prominent midrib
 
                    39. Leaves entire to toothed; petals white or straw to deep purple; cotyledons not doubly folded
ARABIS
                    39' Some leaves lobed; petals pale to lemon yellow; cotyledons doubly folded
DIPLOTAXIS
              35' Fruit ± 4-sided to round in X-section
 
                40. Seeds 2 rows per chamber
RORIPPA
                40' Seeds 1 row per chamber
 
                  41. Petals white to purple
 
                    42. Cauline leaves compound; wet places
Rorippa gambellii
                    42' Cauline leaves ± entire to dissected but not compound; drier places
 
                      43. Plant woody at base; fruit not clearly stalked above receptacle — n White and Inyo Mountains
CAULOSTRAMINA (no key)
                      43' Plant generally not woody at base; fruit ± stalked above receptacle
 
                        44. Fruit stalk < 1 mm, ± as wide as body; fruit often reflexed, not densely clustered; cauline leaves generally not clasping
GUILLENIA
                        44' Fruit stalk < to > 1 mm, narrower than body; fruit not reflexed, often densely clustered; cauline leaves often clasping
THELYPODIUM
                  41' Petals ± yellow (sometimes very pale or pinkish in Guillenia)
 
                    45. Cauline leaves clasping stem
 
                      46. Fruit beak > 5 mm; seed spheric
BRASSICA
                      46' Fruit beak 0 or obscure, any remaining style < 3 mm; seed not spheric
 
                        47. Leaf pinnately lobed to ± compound; plant not glaucous; fruit < 8 cm
BARBAREA
                        47' Leaf ± entire; plant glaucous; fruit 8–13 cm
CONRINGIA (no key)
                    45' Cauline leaves not clasping stem
 
                      48. Fruit beak > 2 mm; seed spheric
BRASSICA
                      48' Fruit beak generally 0, any remaining style < 3 mm; seed not spheric
 
                        49. Fruit ascending to reflexed, 1–7 cm; petals 3–6 mm, white, pale yellow, pinkish
Guillenia lasiophylla
                        49' Fruit erect to ascending, 0.8–10 cm; petals 1.5–10 mm, (pale) yellow
SISYMBRIUM

Key to BRASSICACEAE Group 2

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Leaf generally lobed to compound, sometimes entire
 
    2. Fruit transversely jointed into 2 segments; leaf very fleshy; ocean dunes
CAKILE
    2' Fruit not transversely jointed; leaf not very fleshy; inland
 
      3. Petals white to purplish; fruit ± flat parallel to septum to ± plump; alpine
SMELOWSKIA (no key)
      3' Petals generally yellow (whitish); fruit generally ± cylindric or plump; not alpine
 
        4. Cauline leaf 1–many-pinnately lobed to compound, 1° segments at least again lobed; hairs generally multibranched
DESCURAINIA
        4' Cauline leaf entire to deeply lobed (rarely compound), 1° segments ± entire to toothed; hairs simple or 0
RORIPPA
  1' Leaf entire to shallowly lobed
 
    5. Hairs 0 or simple
 
      6. Lower leaves 3–5 dm; stem 6–20 dm; fruits and seeds generally abortive
ARMORACIA (no key)
      6' Lower leaves < 3 dm; stem generally < 5 dm; fruits and seeds generally fertile
 
        7. Plant aquatic; leaf narrowly awl-like
SUBULARIA (no key)
        7' Plant terrestrial; leaf wider than awl-like
 
          8. Lower petioles >> blades; coastal (North Coast)
COCHLEARIA (no key)
          8' Lower petioles 0 or not > blades; generally not near coast
 
            9. Leaf entire, < 5 mm wide; seed generally 1 per fruit
CUSICKIELLA
            9' Leaf generally toothed or lobed, > 5 mm wide; seeds 2–many per fruit
 
              10. Fruit ± indehiscent, ± flattened or plump; seeds 2–4
CARDARIA
              10' Fruit dehiscent, plump; seeds several–many
RORIPPA
    5' Some hairs generally forked, stellate, or multibranched, especially near stem base (simple in Cardaria)
 
      11. Upper cauline leaves clasping stem
 
        12. Fruit dehiscent, generally > 4-seeded; inflorescence generally not flat-topped, longer than wide
CAMELINA (no key)
        12' Fruit ± indehiscent, 2–4-seeded; inflorescence ± flat-topped, wider than long
CARDARIA
      11' Upper cauline leaves 0 or not clasping stem
 
        13. Herbage hairs mostly stellate; seeds generally 2+ per chamber
 
          14. Fruit not notched
LESQUERELLA
          14' Fruit tip (and often base) ± deeply notched
PHYSARIA (no key)
        13' Herbage hairs mostly simple or multibranched; seed generally 1 per chamber
 
          15. Leaves generally ± basal, < 5 mm wide; fruit not beaked
CUSICKIELLA
          15' Leaves cauline, often > 10 mm wide; fruit beak stout
EUCLIDIUM (no key)

Key to BRASSICACEAE Group 3

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Fruit > 2 × longer than wide, oblong to oblanceolate
ISATIS (no key)2
  1' Fruit ± as long as wide, generally round to elliptic
 
    2. Fruit 2–2.5 mm wide, hooked-hairy, wing 0
ATHYSANUS (no key)
      2. Fruit generally > 3 mm wide, hooked hairs 0, wing obvious, often perforated, radiating rays 0 to very distinct
THYSANOCARPUS

Key to BRASSICACEAE Group 4

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Inflorescence axillary; fruit of 2 nutlet-like halves, wrinkled or ridged, spiny
CORONOPUS
  1' Inflorescence often terminal; fruit not of nutlet-like halves, not wrinkled, ridged, or spiny
 
    2. Seed 1 per chamber
 
      3. Fruit deeply lobed at tip and base between ± round halves; herbage hairs multibranched, stellate
DITHYREA
      3' Fruit sometimes notched at tip but not deeply lobed; herbage hairs 0 or simple
LEPIDIUM
    2' Seeds 2+ per chamber
 
      4. Upper cauline leaves ± clasping stem
 
        5. Fruit obtriangular-obcordate, corners acute; basal leaves generally lobed to deeply dissected
CAPSELLA (no key)
        5' Fruit ± obovate to round, corners rounded; basal leaves entire to dentate
THLASPI
      4' Upper cauline leaves 0 or not clasping stem
 
        6. Leaf deeply lobed to compound, segments linear, often sharp-pointed
POLYCTENIUM (no key)2
        6' Leaf entire to deeply lobed, lobes generally wider than linear, not sharp-pointed
 
          7. Leaf hairs dense, multibranched; petals 15–25 mm; fruit ± 10–20 mm wide
LYROCARPA (no key)
          7' Leaf hairs 0 or sparse, simple or branched; petals < 5 mm; fruit < 5 mm wide
 
            8. Cauline leaves several; filaments scaleless; fruit tip not notched; leaf hairs 0 or branched
HUTCHINSIA (no key)
            8' Cauline leaves 0–few; filaments with a white basal scale; fruit tip shallowly notched; leaf hairs 0 or simple
TEESDALIA (no key)

Key to BRASSICACEAE Group 5

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Fruit 2 × longer than wide
 
    2. Fruit 5–7 mm wide, tip ± tapered, ± sharp-pointed; seed 2.5–3 mm, coat silvery
ANELSONIA (no key)2
    2' Fruit generally < 5 mm wide, tip not tapered or sharp-pointed; seed generally < 2 mm, coat not silvery
DRABA
  1' Fruit < 2 × longer than wide (often ± round)
 
    3. Leaves basal
 
      4. Flowers in racemes; seed wing 0
DRABA
      4' Flowers solitary; seed wing ± 1 mm wide
IDAHOA (no key)
    3' Leaves basal and cauline
 
      5. Flower and fruit > 10 mm wide; fruit stalk above receptacle 7–12 mm
LUNARIA (no key)
      5' Flower and fruit < 5 mm wide; fruit stalk above receptacle 0
 
        6. Inflorescence 1-sided; fruit often ± twisted
HETERODRABA (no key)
        6' Inflorescence not 1-sided; fruit not twisted
 
          7. Leaf hairs forked at base, appressed; fruit valves with prominent midvein
LOBULARIA (no key)
          7' Leaf hairs often multibranched or stellate, often spreading; fruit valves without prominent midvein
 
            8. Style < 1.6 mm
ALYSSUM
            8' Style > 2 mm
LESQUERELLA


Go to the Manual index page for BRASSICACEAE
University & Jepson Herbaria Home Page |

Copyright by the Regents of the University of California