The Jepson Manual: Higher Plants of California

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Names are linked to treatments from the Manual


Key to AGROSTIS

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Rhizomes or stolons well developed, clearly present; perennial herb
 
    2. Plant from stolons
 
      3. Glume back finely scabrous throughout; spikelet falling as 1 unit; inflorescence axes stiff
2 A. semiverticillata
      3' Glume back +/– glabrous, finely scabrous only on keel; floret axis breaking above persistent glumes; inflorescence axes thread-like, flexible
 
        4. Ligule 0.5–2 mm, generally wider than long; stolons < 5 cm; inflorescence widely ovate in outline, 1° branches spreading, spikelets not crowded
3 A. capillaris
        4' Ligule 2–5 mm, longer than wide; stolons 5–100 cm; inflorescence elliptic to lanceolate in outline, 1° branches generally all ascending, spikelets overlapping, crowded
2 A. stolonifera
    2' Plant from rhizomes
 
      5. Floret callus hairs 1.5–2 mm, generally > 1/2 lemma; ligule 4–7 mm
3 A. hallii
      5' Floret callus hairs generally minute, sparse, or 0; ligule generally < 3 mm (except A. gigantea)
 
        6. Palea 0 or minute and << lemma
 
          7. Leaf blade 1–6 mm wide; lemma tip minutely toothed; rhizome < 10 cm; anthers generally > 1 mm
3 A. pallens
          7' Leaf blade < 1 mm wide; lemma tip +/– acute; rhizome < 5 cm; anthers < 1 mm
3 A. variabilis
        6' Palea well developed, +/– 1/2 to slightly < lemma
 
          8. Inflorescence +/– oblong in outline, somewhat dense, branches generally all ascending to erect; montane to alpine, generally > 1500 m
2 A. thurberiana
          8' Inflorescence generally ovate in outline, open, most branches spreading; open, generally disturbed places, generally < 1000 m
 
            9. Rhizomes < 5 cm, slender, not clearly scaly; ligule wider than long; inflorescence branches with spikelets on distal half
3 A. capillaris
            9' Rhizomes < 25 cm, +/– thick, scaly; ligule longer than wide; inflorescence branches with spikelets +/– throughout
2 A. gigantea
  1' Rhizomes or stolons 0; annual or perennial herb
 
    10. Lemma awned from back or near tip
 
      11. Inflorescence open, generally oblong to ovate in outline; spikelets not crowded, inflorescence axes clearly visible
 
        12. Inflorescence generally oblong to lanceolate in outline, 1° branches generally ascending
 
          13. Lemma awned at or above middle; anthers < 0.6 mm; lower leaf sheaths generally glabrous
3 A. exarata
          13' Lemma awned below middle; anthers 1–1.5 mm; lower leaf sheaths finely tomentose
A. hooveri
        12' Inflorescence generally ovate in outline, lowest 1° branches spreading, upper branches generally ascending
 
          14. Lemma back puberulent below middle; floret axis prolonged beyond lemma +/– 1 mm, short-hairy-tufted; palea > 1 mm
A. avenacea
          14' Lemma back glabrous or fine-scabrous; floret axis not prolonged beyond lemma; palea 0 or minute
 
            15. Lemma awned near tip, awn 3–10 mm, wavy; callus hairs < 1/2 lemma, dense; anther 1, persistent in fruit; annual
A. elliottiana
            15' Lemma awned below middle, awn < 2 mm, +/– straight; callus hairs << lemma, sparse; anthers 3, deciduous; perennial herb
2 A. scabra
      11' Inflorescence dense, generally cylindric; spikelets crowded, overlapping, inflorescence axes not clearly visible
 
        16. Primary inflorescence branches generally > 0.5 cm, often evident at base
 
          17. Leaf blade generally < 1 mm wide, +/– inrolled; floret callus glabrous; inflorescence base often partly enclosed by upper leaf; anthers 1–2 mm
2 A. blasdalei
          17' Leaf blade generally 2–10 mm wide, flat; floret callus minutely hairy; inflorescence clearly stalked; anthers +/– 0.5 mm
 
            18. Back of glume fine-scabrous throughout; palea 0.5–0.7 mm; ligule 1.5–2 mm
3 A. densiflora
            18' Back of glume +/– glabrous (keel fine-scabrous); palea +/– 0.3 mm; ligule 2.5–4 mm
3 A. exarata
        16' Primary inflorescence branches generally < 0.5 cm, obscured by densely clustered spikelets
 
          19. Lemma teeth 4, unequal (two < 1 mm, other two 1–1.5 mm); lemma awned below middle; callus densely short-hairy
A. tandilensis
          19' Lemma teeth 0 or 2, equal; lemma awned at or above middle; callus generally sparsely hairy, hairs minute
 
            20. Lemma awned above middle, awn < 3.5 mm, straight; glume tips acute; perennial herb
3 A. densiflora
            20' Lemma awned +/– at middle, awn 3.5–10 mm, generally bent; glume tips narrowly acuminate to awn-like; annual
 
              21. Lemma body 2–4 mm, awn 8–10 mm
A. hendersonii
              21' Lemma body 1.5–2 mm, awn 4–8 mm
A. microphylla
    10' Lemma awnless
 
      22. Spikelets crowded and often overlapping on same branch, spikelet stalks and 2° axes not clearly visible
 
        23. Palea slightly < lemma (or +/– 1/3 × lemma in A. densiflora)
 
          24. Spikelet breaking below glumes, falling as 1 unit; stems often decumbent to long-trailing
2 A. semiverticillata
          24' Floret breaking above persistent glumes; stems generally erect
 
            25. Lower inflorescence branches < 1.5 cm; ligule 1.5–2 mm; anthers ± 0.5 mm
3 A. densiflora
            25' Lower inflorescence branches > 2 cm; ligule 2–5 mm; anthers > 1 mm
2 A. stolonifera
        23' Palea 0 or < 1/3 lemma
 
          26. Leaf blade < 1 mm wide, generally inrolled or folded
 
            27. Anthers > 1 mm; inflorescence branches generally < 0.5 cm; coastal habitats < 100 m
2 A. blasdalei
            27' Anthers < 1 mm; inflorescence branches 0.5–1.5 cm; inland mtns, 1600–4000 m
3 A. variabilis
          26' Leaf blade generally 2–7 mm wide, generally flat
 
            28. Floret callus with hairs 1.5–2 mm, generally slightly > 1/2 lemma
3 A. hallii
            28' Floret callus with hairs 0 or < 0.5 mm, < 1/2 lemma
 
              29. Lower inflorescence branches 1–2 cm; anthers 0.3–0.6 mm
3 A. exarata
              29' Lower inflorescence branches 2–5 cm; anthers 0.7–1.8 mm
3 A. pallens
      22' Spikelets not crowded, generally well spaced on same branch, axes clearly visible
 
        30. Primary inflorescence axes branched 1–2 × above middle, spikelets 0 on lower 1/2
 
          31. Leaf blades 2–4 mm wide; glumes 2–3 mm; palea 1/4–1/5 lemma; anthers 0.6–0.8 mm; moist areas
A. oregonensis
          31' Leaf blades 0.7–2 mm wide; glumes 1.5–2.7 mm; palea 0 or minute, << lemma; anthers < 0.7 mm; well drained areas
 
            32. Leaves basal and cauline; inflorescence +/– 2 × longer than wide; 1° axes +/– stiff
A. idahoensis
            32' Leaves mostly basal; inflorescence +/– long as wide; 1° axes flexible, lower +/– arched
2 A. scabra
        30' Primary inflorescence axes branched 1–2 × from base upwards, spikelets distributed throughout
 
          33. Floret callus with hairs 1.5–2 mm, generally slightly > 1/2 lemma; anthers > 1.5 mm
3 A. hallii
          33' Floret callus glabrous or with minute hairs << lemma; anthers generally < 1.5 mm (except some A. pallens)
 
            34. Palea 0 or minute, << lemma
 
              35. Leaf blades 1–6 mm wide; plants generally 10–70 cm; anthers 0.7–1.8 mm
3 A. pallens
              35' Leaf blades generally < 1 mm wide; plants 4–30 cm; anthers 0.5–0.7 mm
3 A. variabilis
            34' Palea > 1/2 lemma
 
              36. Inflorescence branches spreading to ascending; anthers 1–1.5 mm; wet or disturbed areas, generally < 1500 m
 
                37. Ligule 0.5–2 mm, generally wider than long; inflorescence branches with spikelets on distal half
3 A. capillaris
                37' Ligule 2–6 mm, longer than wide; inflorescence branches with spikelets +/– throughout
2 A. gigantea
              36' Inflorescence branches ascending to erect; anthers < 1 mm; moist to +/– dry areas, generally > 1500 m
 
                38. Inflorescence 2–5 cm, slightly flattened; plants generally < 15 cm; alpine
A. humilis
                38' Inflorescence generally 4–11 cm, not clearly flattened; plants 8–50 cm; < subalpine
2 A. thurberiana


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