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SCROPHULARIACEAE

FIGWORT FAMILY

Lawrence R. Heckard, Family Coordinator

Annual to shrubs, generally glandular, some green root-parasites
Stem generally round
Leaves generally alternate, simple, generally ± entire; stipules generally 0
Inflorescence: spike to panicle, generally bracted, or flowers 1–2 in axils
Flower bisexual; calyx lobes generally 5; corolla generally strongly bilateral, generally 2-lipped (upper lip generally 2-lobed, lower lip generally 3-lobed); stamens generally 4 in 2 pairs, generally included, a 5th (generally uppermost) sometimes present as a staminode; pistil 1, ovary superior, chambers generally 2, placentas axile, style 1, stigma lobes generally 2
Fruit: capsule, generally ± ovoid, loculicidal or septicidal
Seed: coat sculpture often characteristic
Genera in family: ± 200 genera, 3000 species: ± worldwide; some cultivated as ornamental (e.g., Antirrhinum, Mimulus, Penstemon ) or medicinal (Digitalis )
Recent taxonomic note: Recently treated to include only Buddleja, Scrophularia, and Verbascum in CA; other genera moved to Orobanchaceae (Castilleja, Cordylanthus, Orthocarpus, Pedicularis, Triphysaria), Phrymaceae (Mimulus), and Plantaginaceae (= Veronicaceae sensu Olmstead et al.)
Key to genera by Elizabeth Chase Neese & Margriet Wetherwax.

VERONICA

SPEEDWELL, BROOKLIME

Margriet Wetherwax

Annual, perennial herb
Stem erect or prostrate
Leaves opposite
Inflorescence: raceme, terminal or axillary, or flowers solitary in axils; bracts small, alternate
Flower: sepals generally 4(5), ± free, generally unequal; corolla ± rotate, 4-lobed, upper lobe wide (formed by fusion of upper pair), blue or violet to white; stamens 2, exserted; stigma head-like
Fruit: capsule, flattened perpendicular to septum, generally obcordate, loculicidal and septicidal
Species in genus: ± 250 species: n temp, especially Eurasia
Etymology: (Possibly named for Saint Veronica)

Native

V. serpyllifolia L. subsp. humifusa (Dickson) Syme

Perennial, rhizomed, ± hairy
Stem erect at tips only, 5–30 cm
Leaf 10–25 mm, elliptic to widely ovate, obtuse, entire to crenate
Inflorescence terminal, glandular-hairy; pedicels 2.5–7 mm
Flower: sepals 2.5–4 mm, ± equal, oblong to ovate; corolla 6–7 mm, bright blue; style 2–3 mm
Fruit 2.8–3.7 mm, wider than long, ± glandular-hairy; notch 0.3–0.8 mm
Seed 0.5 mm, flat
Chromosomes: 2n=14,28
Ecology: Moist streambanks, lakeshores, meadows
Elevation: 1700–3000 m.
Bioregional distribution: Cascade Range, Sierra Nevada, San Bernardino Mountains, East of Sierra Nevada
Distribution outside California: to Alaska, ne US, New Mexico; also in S.America, Eurasia
Synonyms: var. h. Dickson
Ssp. s. less hairy, pedicels shorter, flowers and fruit smaller; uncommon lawn weed in w CA-FP; native to Eur.

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bioregional map for VERONICA%20serpyllifolia%20subsp.%20humifusa being generated
 


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