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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

SCROPHULARIACEAE

FIGWORT FAMILY

Lawrence R. Heckard, Family Coordinator

Annual to shrubs, generally glandular, some green root-parasites
Stem generally round
Leaves generally alternate, simple, generally ± entire; stipules generally 0
Inflorescence: spike to panicle, generally bracted, or flowers 1–2 in axils
Flower bisexual; calyx lobes generally 5; corolla generally strongly bilateral, generally 2-lipped (upper lip generally 2-lobed, lower lip generally 3-lobed); stamens generally 4 in 2 pairs, generally included, a 5th (generally uppermost) sometimes present as a staminode; pistil 1, ovary superior, chambers generally 2, placentas axile, style 1, stigma lobes generally 2
Fruit: capsule, generally ± ovoid, loculicidal or septicidal
Seed: coat sculpture often characteristic
Genera in family: ± 200 genera, 3000 species: ± worldwide; some cultivated as ornamental (e.g., Antirrhinum, Mimulus, Penstemon ) or medicinal (Digitalis )
Recent taxonomic note: Recently treated to include only Buddleja, Scrophularia, and Verbascum in CA; other genera moved to Orobanchaceae (Castilleja, Cordylanthus, Orthocarpus, Pedicularis, Triphysaria), Phrymaceae (Mimulus), and Plantaginaceae (= Veronicaceae sensu Olmstead et al.)
Key to genera by Elizabeth Chase Neese & Margriet Wetherwax.

VERONICA

SPEEDWELL, BROOKLIME

Margriet Wetherwax

Annual, perennial herb
Stem erect or prostrate
Leaves opposite
Inflorescence: raceme, terminal or axillary, or flowers solitary in axils; bracts small, alternate
Flower: sepals generally 4(5), ± free, generally unequal; corolla ± rotate, 4-lobed, upper lobe wide (formed by fusion of upper pair), blue or violet to white; stamens 2, exserted; stigma head-like
Fruit: capsule, flattened perpendicular to septum, generally obcordate, loculicidal and septicidal
Species in genus: ± 250 species: n temp, especially Eurasia
Etymology: (Possibly named for Saint Veronica)

Native

V. scutellata L.

MARSH SPEEDWELL

Perennial, rhizomed, ± glabrous
Stem decumbent to erect, slender, 10–60 cm, generally glabrous
Leaf sessile, 20–40 mm, linear-lanceolate, ± entire, often purple-tinged
Inflorescences alternate, open; flowers 5–20; bracts < 1/2 pedicels, linear; pedicels 6–16 mm, spreading at right angles in fruit
Flower: sepals ± 3 mm, ± equal, ovate; corolla 5–7 mm, white or blue, purple-lined
Fruit 3–4 mm, longer than wide; notch 0.4–1 mm
Seed ± 1.5 mm, flat
Chromosomes: 2n=18
Ecology: Wet meadows, ponds
Elevation: < 2200 m.
Bioregional distribution: Cascade Range, Sierra Nevada, San Bernardino Mountains
Distribution outside California: to British Columbia, e US; Europe

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bioregional map for VERONICA%20scutellata being generated
 


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