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BRASSICACEAE

MUSTARD FAMILY

Reed C. Rollins, except as specified

Annual to subshrub
Leaves generally basal and cauline, alternate, generally simple; stipules 0
Inflorescence: generally raceme
Flower bisexual; sepals 4, free; petals (0)4, free, generally white or yellow, often clawed; stamens generally (2,4)6, generally 4 long, 2 short; ovary 1, superior, chambers generally 2, septum membranous, connecting 2 parietal placentas, style 1, stigma simple or 2-lobed
Fruit: generally capsule ("silique") with 2 deciduous valves, sometimes breaking transversely or indehiscent
Seeds 1–many per chamber
Genera in family: 300+ genera, 3000+ species: worldwide, especially cool regions; some cultivated for food (especially Brassica, Raphanus ) and ornamental
Recent taxonomic note: Recently treated to include Capparaceae [Rodman et al. 1993 Ann Missouri Bot Gard 80:686–699; Rollins 1993 Cruciferae of Continental North America. Stanford Univ Press]
Family description, key to genera by Robert A. Price.

STREPTANTHUS

JEWELFLOWER

Roy E. Buck, Dean W. Taylor, and Arthur R. Kruckeberg

Annual to perennial herb, glabrous to bristly, generally ± glaucous
Leaves ± entire to pinnately compound; basal generally rosetted, generally ± petioled; cauline linear to (ob)ovate, often clasping
Inflorescence generally ± open; bracts generally 0
Flower biradial or bilateral; calyx generally ± urn-shaped, sepals erect, generally not green, bases ± pouch-like, generally keeled; petals generally exserted, blade generally narrower than claw, ± channeled, margins ± wavy, generally ± scarious; stamens generally in 3 free pairs; style 0 or short, stigma generally ± entire, blunt
Fruit long, generally strongly compressed parallel to septum
Seeds generally compressed, generally ± winged
Species in genus: ± 40 species: sw US, n Mex
Etymology: (Greek: twisted flower, from wavy-margined petals)
Reference: [Dolan & LaPré 1989 Madroño 36:33–40; Kruckeberg & Morrison 1983 Madroño 30:230–244]
Caulanthus sometimes including here. Calluses on leaf margins of some mimic pierid butterfly eggs, reducing larval herbivory. Variable, complex; needs study.

Native

S. glandulosus Hook.

JEWELFLOWER

Annual 1–10 dm, simple to branched throughout, generally ± densely bristly, especially below
Leaves < 13 cm; basal ± oblanceolate, coarsely dentate to ± lobed, narrowed to ± winged petiole; cauline ± linear to (ob)lanceolate, entire to ± lobed
Inflorescence ± straight; pedicels generally < sepals
Flower: calyx bilateral, sepals 4–9 mm, glabrous or sparsely hairy; petals 6–15 mm, upper >, generally darker than lower; upper filament pair exserted, fused at least in basal 2/3, recurved, with reduced ± sterile anthers
Fruit spreading to ± erect (less commonly deflexed), 3–11 cm, straight to ± curved
Chromosomes: 2n=28
Ecology: Dry, open grasslands, chaparral, open conifer/oak woodland, sometimes on serpentine, mostly away from coast
Elevation: 15–1300 m.
Bioregional distribution: s North Coast Ranges, San Francisco Bay Area, n&c South Coast Ranges
Distribution outside California: sw Oregon?

Native

subsp. pulchellus (Greene) Kruckeb.

MOUNT TAMALPAIS JEWELFLOWER


Stem generally 1–3 dm
Inflorescence dense, ± 1-sided
Leaves < 6 cm
Flower: sepals 4–7 mm, reddish purple; petals 8–13 mm, ± purple
Ecology: Habitats ± of sp.
Elevation: 150–800 m.
Bioregional distribution: nw San Francisco Bay Area (Marin Co.)
Synonyms: var. p. (Greene) Jeps

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