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©Copyright 1993 by the Regents of the University of California
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ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

SOLIDAGO

GOLDENROD

John C. Semple

Perennial from woody caudex or rhizome, branched above
Leaves alternate, resinous, often sessile
Inflorescence: heads radiate, few–many, in ± flat-topped to panicle-like, often ± 1-sided clusters; involucre cylindric to bell-shaped (wider when dry); phyllaries in 3–5 graduated, overlapping series, midrib generally ± swollen, translucent
Ray flowers few–many; ligules yellow
Disk flowers few–many; corollas yellow, generally glabrous; style branches finely papillate, appendage triangular
Fruit obconic, compressed; pappus of 25–45 long-barbed bristles in 1 series
Species in genus: ± 150 species: especially North America (South America, Eurasia)
Etymology: (Greek: make-well, from purported medicinal value)
Reference: [Semple et al. 1990 Can J Bot 68:2070–2082]

Native

S. gigantea Aiton

SMOOTH GOLDENROD

Herbage glabrous to ± strigose
Stems < 15 dm, rhizomed, ± waxy
Leaves: middle cauline largest, 5–15 cm, narrowly elliptic to ± lanceolate, 3-veined, toothed
Inflorescence ± open, pyramid-shaped; heads many; involucre 3–4 mm; phyllaries generally lanceolate, outer 1/4–1/3 length inner
Ray flowers 8–15; ligules 2–2.5 mm
Disk flowers 7–12; corollas 3–4 mm
Fruit 1–1.5 mm, sparsely strigose
Chromosomes: 2n=18,36,54
Ecology: Moist streambanks, lakesides
Elevation: 1000–1500 m.
Bioregional distribution: n High Sierra Nevada, Modoc Plateau
Distribution outside California: to e N.America
Like S. canadensis.

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