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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

SOLIDAGO

GOLDENROD

John C. Semple

Perennial from woody caudex or rhizome, branched above
Leaves alternate, resinous, often sessile
Inflorescence: heads radiate, few–many, in ± flat-topped to panicle-like, often ± 1-sided clusters; involucre cylindric to bell-shaped (wider when dry); phyllaries in 3–5 graduated, overlapping series, midrib generally ± swollen, translucent
Ray flowers few–many; ligules yellow
Disk flowers few–many; corollas yellow, generally glabrous; style branches finely papillate, appendage triangular
Fruit obconic, compressed; pappus of 25–45 long-barbed bristles in 1 series
Species in genus: ± 150 species: especially North America (South America, Eurasia)
Etymology: (Greek: make-well, from purported medicinal value)
Reference: [Semple et al. 1990 Can J Bot 68:2070–2082]

Native

S. californica Nutt.

CALIFORNIA GOLDENROD

Herbage generally ± densely short-soft-hairy
Stems 2–15 dm, short-rhizomed
Leaves: lower < 14 cm, oblanceolate to obovate, serrate, sometimes 3-veined, base tapered; upper much reduced
Inflorescence long, 1-sided, pyramid- or wand-shaped, many-headed (or short, raceme-like, few-headed); involucre 3–5 mm; phyllaries generally narrow, acute, strigose, outer 1/4–1/3 length inner
Ray flowers 6–11; ligules 3–5 mm
Disk flowers 6–17; corollas 3–5 mm
Fruit 0.7–1.5 mm, ± densely strigose
Chromosomes: 2n=18,36
Ecology: Woodland margins, grassland, disturbed soils
Elevation: < 2300 m.
Bioregional distribution: California Floristic Province, Modoc Plateau, White and Inyo Mountains
Distribution outside California: Oregon, Baja California
Flowering time: Jul–Oct
Horticultural information: SUN: 4, 5, 6 &IRR: 1, 2, 3, 15, 16, 17, 18, 19, 20, 21, 22, 23 &SHD: 7, 8, 9, 14.