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SAXIFRAGACEAE

SAXIFRAGE FAMILY

Patrick E. Elvander

Perennial or subshrub from caudex or rhizome, generally ± hairy
Stem often ± leafy on lower half, rarely trailing and leafy throughout
Leaves generally simple, basal or sometimes cauline, generally alternate, generally petioled; veins ± palmate
Inflorescence: panicle, generally ± scapose
Flower generally bisexual, generally radial; hypanthium free to ± fused to ovary; calyx lobes generally 5; petals generally 5, free, generally clawed, generally white; stamens generally 5 or 10; pistils 2 and simple or 1 and compound (chambers 1–2, placentas 2–4, axile or parietal), ovary superior to inferior, sometimes more superior in fruit, styles generally 2
Fruit: 2 follicles or 2–4-valved capsule
Seeds generally many, small
Genera in family: 40 genera, 600 species: especially n temp, arctic, alpine; some cultivated (Bergenia , Darmera , Heuchera , Saxifraga , Tellima , Tolmiea )
Reference: [Soltis 1988 Syst Bot 13:64–72]

SAXIFRAGA

SAXIFRAGE

Plant generally ± hairy, often glandular; caudex or rhizome generally not woody, generally scaly
Leaf: blade oblanceolate to round, base tapered to reniform, margin entire or toothed
Inflorescence: flowers 1–many; bracts scale-like
Flower: hypanthium free or ± fused to ovary; petals sometimes spotted; stamens 10, filaments generally flat; pistil 1 (chambers 2, placentas 2, axile) or 2 (each with 1 chamber, 1 marginal placenta), ovary superior to ± inferior (sometimes more superior in fruit), styles 2, free throughout
Fruit: capsule or 2 follicles
Species in genus: ± 400 species: especially cool n temp
Etymology: (Latin: rock-breaking)
Reference: [Elvander 1984 Syst Bot Monogr 3:1–44]

Introduced

S. stolonifera Meerb.

STRAWBERRY-GERANIUM

Caudex branched, generally producing stolons; bulblets 0
Leaf 8–14 cm; petiole 5–8 cm; blade ± round, base shallowly cordate to truncate, teeth irregular, ± round
Inflorescence ± 3–4 dm, open
Flower bilateral; sepals reflexed, < petals, ovate; petals unequal, lower two 10–15 mm, >> upper 3 generally with 1–3 spots; filaments generally widened toward tips; nectaries band-like; ovary superior
Fruit: ± follicles
Chromosomes: 2n=36,54
Ecology: Moist, shady places
Elevation: < 200 m.
Bioregional distribution: s Outer North Coast Ranges, expected more widely
Distribution outside California: native to China, Japan
Synonyms: S. sarmentosa L
Cult as ornamental.

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