This page is based on the 1993 Jepson Manual.
Please see the Jepson eFlora for up-to-date information about California vascular plants.
|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL||
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
Print edition is available from the University of California Press
|The second edition of The Jepson Manual (2012) is available from the University of California Press|
|See also the Jepson eFlora, which parallels the Second Edition|
Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5many, pistils (0)1many, simple or compound; ovary superior to inferior, styles 15
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 15
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303332,344401,611662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.
Perennial to shrub, often bramble-forming, often prickly, prostrate or clambering to erect
Leaves generally palmately lobed to compound; leaflets often stalked, toothed
Flower: hypanthium shallow; bractlets 0; sepals 5, generally reflexed, tips generally linear; petals 5; stamens generally > 20; pistils fewmany, ovaries superior, jointed to slender to club-shaped styles
Fruit: aggregate of sweet, fleshy-coated achenes (drupelets) that generally separate jointly from receptacle (raspberry-like) or separate jointly with part of fleshy receptacle (blackberry-like)
Species in genus: 200700 species: worldwide, especially n temp, Andes
Etymology: (Latin: ancient name for bramble)
Coastal forms often have smaller, hairier, rounder leaves; hybrids and other escapes from cultivated expected.
Arched to erect brambles
Stem 410 mm diam, round; prickles many, slender to wide-based, straight to ± curved (especially in inflorescence)
Leaf compound; stipules linear; petiole ± 15 cm; leaflets 3(7), ± ovate, shallowly lobed, irregularly toothed, white below; longest leaflet stalk ± 520 mm; longest leaflet blade ± 26 cm
Inflorescence: ± cyme, few-flowered, glandular or not
Flower: sepal tips 13 mm; petals 35 mm, oblanceolate-elliptic, white; pistils generally > 15
Fruit raspberry-like, ± spheric, red-purple to black, puberulent
Ecology: Generally ± rocky, especially moist areas
Elevation: 402400 m.
Bioregional distribution: California Floristic Province (except coast, Great Central Valley)
Distribution outside California: to British Columbia, Montana, Nevada
Glandular plants (TR, PR) have been called var. bernardinus (Greene) Jeps.; simple-lvd plants (KR) have been called var. trinitatis A. Berger
Horticultural information: 4, 5, 6 &IRR: 1, 2, 7, 8, 9, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24.
|YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).|