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PLANTAGINACEAE

PLANTAIN FAMILY

Lauramay T. Dempster

Annual, perennial herb, generally scapose
Leaves generally basal, simple, with longitudinal ribs; stipules 0
Inflorescence: spike, generally terminal, generally dense; flowers few–many, each subtended by 1 bract
Flower unisexual or bisexual, generally radial; calyx deeply 4-lobed, lobes generally overlapping, persistent, margin mostly scarious; corolla salverform or cylindric, lobes 4, spreading to erect, scarious, persistent, colorless; stamens 2–4, alternate corolla lobes, epipetalous; ovary superior, 2–4-chambered, ovules several per chamber, style 1, stigma long, hairy
Genera in family: 3 genera, ± 270 species: worldwide, especially temp; some weedy, some (especially P. afra , psyllium) cultivated for laxative
Recent taxonomic note: Recently treated to include Callitrichaceae, and most non-parasitic CA genera of Scrophulariaceae (but not Mimulus, Scrophularia, Verbascum); = Veronicaceae sensu Olmstead et al. [Angiosperm Phylogeny Group 1998 Ann Missouri Bot Gard 85:531–553; Olmstead et al. 2001 Mol Phylog Evol 16:96–112]

PLANTAGO

PLANTAIN

Annual, perennial herb
Stem decumbent to erect
Flower generally bisexual; corolla radial, sometimes bilateral
Fruit: capsule, circumscissile ± at or below middle
Seeds 2–many, gelatinous when wet
Species in genus: ± 250 species: worldwide
Etymology: (Latin: sole of foot)

Introduced

P. lanceolata L.

ENGLISH PLANTAIN

Perennial from stout caudex, taprooted, short-hairy
Leaf 5–25 cm, lanceolate to lance-oblong, tapered gradually to petiole; margin generally finely dentate
Inflorescences few–many, 20–80 cm including peduncle; spike 2–8 cm, ± ovoid, becoming cylindric, dense; bract ± = sepals, widely ovate, acute, not exserted
Flower: corolla lobes 2–2.5 mm, spreading, ovate, tips ± acute; stamens 4
Seeds 1–2, ± 3 mm
Chromosomes: 2n=12,24,96
Ecology: Common. Weed of waste places, lawns, roadsides
Elevation: < 1600 m.
Bioregional distribution: California Floristic Province
Distribution outside California: native to Europe

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