Jepson Flora Project: Jepson Interchange    

link to manual TREATMENT FROM THE JEPSON MANUAL (1993) previous taxon | next taxon
Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora.



Alan R. Smith and Thomas Lemieux

Perennial, in soil or on or among rocks; rhizome creeping to erect, scaly
Leaves generally all ± alike (or of 2 kinds, fertile and sterile), generally < 50 cm, often < 25 cm; petiole generally thin, wiry, often dark, in transverse section with vascular strands generally 1–3, less often many in a circle; blade generally pinnate or ± palmate-pinnate (see Adiantum ), often 2 or more compound, lower surface often with glands, ± powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or otherwise, veins generally free
Sporangia in sori or not, marginal, submarginal, or along veins, sometimes covered by recurved, often modified segment margins (false indusia); true indusia 0; spores spheric, sides sometimes flat, scar with 3 radiating branches
Genera in family: ± 40 genera, 500 species: worldwide, especially dry areas. Definition of Cheilanthes and related genera problematic; traditional limits often untenable.



Plant in soil or rock crevices; rhizome short-creeping-decumbent, generally 3–5(8) mm wide, scales linear-lanceolate, with darker mid-stripe
Leaf: petiole 5–20(32) cm, 0.5–2(3) mm wide; blade generally 2–3-pinnate, 2–8(15) cm, triangular or generally 5-sided, with white or yellow exudate on lower surface, with or without exudate on upper surface, main axis shallowly to deeply grooved on upper surface; lowermost 1° leaflets each strongly asymmetric (more developed on basal surface); veins free
Sporangia along veins ± throughout; segment margins unmodified, recurved or not
Species in genus: 2 species: w North America. A puzzling complex of intergrading chemical, chromosomal, and morphological variants (see Yatskievych et al. 1990 Amer Fern J 80:9–17); for these we prefer the rank of var. but have used subsp. here because combinations at that rank already exist
Horticultural information: DFCLT.


P. triangularis (Kaulf.) Yatsk., Windham & E. Wollenw.

Rhizome tip, scales without exudate
Leaf: petiole brown to reddish brown, with or without exudate; blade 3–10(18) cm, generally pale to dark green, upper surface generally without exudate
Chromosomes: 2n=60,90,120,150
Ecology: Common. Generally shaded slopes or rocky areas
Elevation: < 2300 m.
Bioregional distribution: California Floristic Province, Modoc Plateau (caves in Lava Beds National Monument), East of Sierra Nevada, Desert Mountains
Distribution outside California: to British Columbia, Nevada, Arizona, Baja California
Synonyms: Pityrogramma t. (Kaulf.) Maxon


subsp. maxonii (Weath.) Yatsk., Windham & E. Wollenw.

Leaf: blade upper surface generally with sparse, minute (0.1 mm), yellowish glands, lower surface often with many yellowish or reddish glands, margins not recurved; lower 1° leaflets 2–5(7) cm; upper 1° leaflets and 2° leaflets on basal surface of lowermost 1° leaflets deeply pinnately lobed to nearly 1-pinnate
Ecology: Generally ± shaded, near rocks, boulders
Elevation: 300–1400 m.
Bioregional distribution: San Bernardino Mountains, Peninsular Ranges, Desert Mountains
Distribution outside California: Arizona, Baja California
Synonyms: Pityrogramma t. (Kaulf.) Maxon var. m. Weath

previous taxon | next taxon
bioregional map for PENTAGRAMMA%20triangularis%20subsp.%20maxonii being generated
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

Retrieve Jepson Interchange Index to Plant Names entry for Pentagramma triangularis subsp. maxonii
Retrieve dichotomous key for Pentagramma
Overlay Consortium of California Herbaria specimen data by county on this map
Show other taxa with the same California distribution | Read about bioregions | Get lists of plants in a bioregion
Return to the Jepson Interchange main page
Return to treatment index page
  • This page is no longer being maintained.

University & Jepson Herbaria Home Page |
General Information | University Herbarium | Jepson Herbarium |
Visiting the Herbaria | On-line Resources | Research |
Education | Related Sites
Copyright © by the Regents of the University of California