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ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

MADIA

TARWEED

Annual or perennial herb, generally densely glandular, aromatic
Stems 1–several, generally simple below, ± branched above
Leaves generally opposite below, alternate above, generally linear to lanceolate, entire to slightly toothed
Inflorescence: heads generally radiate, generally peduncled, few–many; phyllaries generally 1–20, free, enclosing (and falling with) ray achenes; receptacle ± flat, generally glabrous; chaff scales generally ± fused, in ring between ray and disk flowers
Ray flowers generally 1–20, sometimes minute; ligules 2–3-lobed, generally yellow
Disk flowers 1–many, sometimes staminate; corollas yellow or maroon; anther tips triangular-ovate; style tips linear to oblong, acute, bristly
Fruit club-shaped or obovoid; ray achenes compressed, thickened, or 3-angled (1 angle toward center of head), ridged, sometimes beaked; pappus 0 or of short scales; disk achenes ± symmetric; pappus 0 or of 4–10 scales or bristles
Species in genus: 21 species: w North America, sw South America
Recent taxonomic note: *See revised taxonomy of Baldwin 1999 Novon 9:462–471.
Etymology: (Chilean name)
Reference: [Nelson & Nelson 1980 Brittonia 32:323–325]

Native

M. rammii Greene

Annual 1–6 dm
Stems: branches ascending, moderately leafy, soft-hairy below, densely stalked-glandular above; glands golden
Leaves 1.5–5 cm, mostly scattered, strigose to bristly
Inflorescence: heads generally in open cymes; peduncles 3–10 cm; involucre 3.8–4.8 mm, widely urn-shaped; phyllaries acute, soft-hairy to short-bristly, not glandular, tips not flat
Ray flowers 15–20; tubes 0.5–1 mm, ligules 4–9 mm, bright yellow, sometimes ± purple- or brown-lined
Disk flowers 20–65, staminate; corollas 3–4 mm; anthers black
Fruit: ray achenes 2.4–3.5 mm, strongly bowed out, compressed side-to-side, beaked, pappus 0 or scales minute, fringed; disk ovaries linear, pappus scales 5–7, 3–4 mm, linear, bristle-like, ± bent, ciliate, purplish
Chromosomes: 2n=16
Ecology: Grassy slopes
Elevation: 400–1600(2700) m.
Bioregional distribution: n Sierra Nevada
Recent taxonomic note: *Jensia rammii (Greene) B.G. Baldwin

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