Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
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Dieter H. Wilken, except as specifed

Annual, perennial herb, shrub, glabrous to hairy, generally aromatic
Stems generally erect, generally 4-angled
Leaves generally simple to deeply lobed, opposite, generally gland-dotted
Inflorescence: cyme, generally clustered around stem, head-like, separated by evident internodes (terminal in Monardella ) or collectively crowded, spike-like to panicle-like (sometimes raceme or flowers 2–12); subtended by leaves or bracts; flowers sessile or pedicelled
Flower generally bisexual; calyx generally 5-lobed, radial to bilateral; corolla generally bilateral, 1–2-lipped, upper lip entire or 2-lobed, ± flat to hood-like, sometimes 0, lower lip generally 3-lobed; stamens generally 4, generally exserted, paired, pairs unequal, sometimes 2, staminodes 2 or 0; ovary superior, generally 4-lobed to base, chambers 2, ovules 2 per chamber, style 1, arising from center at junction of lobes, stigmas generally 2
Fruit: nutlets 4, generally ovoid to oblong, smooth
Genera in family: ± 200 genera, 5500 species: worldwide. Many cultivated for herbs, oils (Lavandula , lavender; Mentha , mint; Ocimum , basil; Rosmarinus , rosemary; Thymus , thyme), some cultivated as ornamental (in CA Cedronella , Leonotis , Phlomis )
Reference: [Cantino & Sanders 1986 Syst Bot 11:163–185]



Perennial from rhizomes, glabrous or hairy
Stems erect, branched or not
Leaves short-petioled to sessile, generally ovate to lanceolate; margin toothed to deeply lobed or cut below middle
Inflorescences axillary, each head-like, subtended by leaves
Flower: calyx generally 5-lobed, lobes ± equal, obtuse to short-awned; corolla slightly bilateral, not 2-lipped, generally 4-lobed, lobes ± unequal, odd lobe notched or entire; stamens 2, exserted, staminodes 2, minute, club-shaped; style exserted
Fruit: nutlets ± compressed, edge corky-thickened, truncate or rounded
Species in genus: 14 species: temp North America, Eurasia, 1 species in Australia
Etymology: (Greek: wolf foot, from French common name)
Reference: [Henderson 1962 Amer Midl Naturalist 68:95–135]


L. asper Greene

Rhizomes thicker and tuber-like near tip
Stems erect, 3–8(10) dm, puberulent to short-hairy
Leaf 2.5–7(9) cm, ± sessile, lanceolate to narrowly elliptic, serrate, glabrous to puberulent
Flower: calyx lobes awl-like, acuminate to short-awned; corolla 3–5 mm, slightly > calyx, white
Fruit: nutlet 1.5–2 mm; top truncate, sometimes minutely toothed
Chromosomes: 2n=22
Ecology: Uncommon. Moist areas, marshes, streambanks
Elevation: < 1300 m.
Bioregional distribution: Deltaic Great Central Valley, San Francisco Bay Area, Great Basin Floristic Province
Distribution outside California: to w Canada, Great Plains
Flowering time: Jun–Oct
Synonyms: L. lucidus Benth. misapplied

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bioregional map for LYCOPUS%20asper being generated
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

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