ROSACEAE

ROSE FAMILY

Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5–many, pistils (0)1–many, simple or compound; ovary superior to inferior, styles 1–5
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 1–5
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303–332,344–401,611–662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.

HORKELIA

Barbara Ertter

Perennial, generally ± glandular, generally resinous-smelling; caudex generally branched
Stem generally ascending to erect
Leaves generally basal, odd-1-pinnate, generally ± flat; cauline alternate, reduced upward; uppermost lateral leaflets generally ± fused with terminal
Inflorescence: cyme, open or of dense clusters; pedicels generally straight
Flower: hypanthium a ± flat-bottomed cup, width ± 2 X length; bractlets 5, generally 2/3 sepals; sepals 5, often reflexed; petals 5, generally ± = sepals, blunt, white; stamens 10, filaments flat, often forming a tube; pistils 2–many, ovaries superior, styles jointed below fruit tip, ± thicker at base
Fruit: achene
Species in genus: 19 species: w North America
Etymology: (J. Horkel, German plant physiologist, 1769–1846)
Data apply to basal leaves, pressed hypanthia.

Native

H. yadonii Ertter

SANTA LUCIA HORKELIA

Plant tufted to matted, grayish green
Stem generally 20–60 cm; hairs spreading
Leaf generally 6–20 cm; leaflets 7–16 per side, separated to ± crowded, generally 4–15 mm, wedge-shaped to round, ± 3–20-toothed or -lobed, often deeply notched, hairs generally dense
Inflorescence ± open, generally 5–10-flowered; pedicels generally 1–7 mm
Flower: hypanthium width generally 3–6 mm, ± > 2 X length, inner wall ± hairy; bractlets 1–2 mm wide, lanceolate to ovate; sepals generally 4–6 mm; petals 3–5 mm, oblanceolate to elliptic; filaments 1–2 mm, bases 0.5–1 mm wide, anthers ± 1 mm; pistils generally > 20, styles 2.5–3 mm
Fruit ± 1.5 mm
Ecology: UNCOMMON. Sandy meadow edges, seasonal streambeds in chaparral or foothill-pine woodland
Elevation: 350–1900 m.
Bioregional distribution: Outer South Coast Ranges.Sometimes confused with H. cuneata subsp. sericea , H. rydbergii , or H. tenuiloba
Horticultural information: IRR: 7, 8, 9, 14, 15, 16, 17, 19, 20, 21, 22, 23, 24; GRCVR.