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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Ronald L. Hartman (except Silene)

Annual, biennial, perennial herb, rarely dioecious, taprooted or rhizome generally slender
Leaves simple, generally opposite; stipules generally 0; petiole generally 0; blade entire, sheath generally 0
Inflorescence: cyme, generally open; flowers few–many or flower solitary and axillary; involucre generally 0
Flower generally bisexual, radial; hypanthium sometimes present; sepals generally 5, ± free or fused into a tube, tube generally herbaceous between lobes or teeth; awns generally 0; petals generally 5 or 0, generally tapered to base (or with claw long, blade expanded), entire to 2–several-lobed, blade generally without scale-like appendages (inner surface), generally without ear-like lobes at base; stamens generally 10, generally fertile, generally free, generally from ovary base; nectaries generally 0; ovary superior, generally 1-chambered, placentas basal or free-central, styles 2–5 or 1 and 2–3-branched
Fruit: capsule or utricle (rarely modified, dehiscent), generally sessile
Seeds: appendage generally 0
Genera in family: 85 genera, 2400 species: widespread, especially arctic, alpine, temp, n hemisphere; some cultivated (Agrostemma, Arenaria, Cerastium, Dianthus, Gypsophila, Lychnis, Saponaria, Silene, Vaccaria ).



Annual, ascending to erect, taprooted
Leaves: basal blades linear to oblanceolate, short-petioled; cauline blades narrowly oblanceolate to oblong-ovate, sessile; vein 1
Inflorescence: cyme, umbel-like, terminal; flowers 3–12; involucral bracts 5–15, ± 0.3–1.8 mm; pedicels 0.5–2.5 cm
Flower: sepals 5, 3–4.5 cm, lanceolate to ovate, glabrous; petals 5, 3.3–5 mm, generally minutely toothed; stamens 5–10; styles 3, 0.5–1.4 mm
Fruit: capsule, ovoid or ovoid-cylindric, straight; teeth 6, recurved
Seeds many, reddish brown
Species in genus: 6 species: temp Eurasia
Etymology: (Greek: all bone, humorous reference to frailty of plant)
Reference: [Shinners 1965 Sida 2:119–128]


H. umbellatum L. subsp. umbellatum

Plant 5–25 cm
Stems 1–many, unbranched, glandular-hairy near middle
Leaves: cauline 1–4 pairs in lower 1/2; blades 2–25 mm, glabrous or margin glandular-hairy
Inflorescence: bracts whorled, 1–2 mm, partly scarious; pedicels ascending to erect, reflexed in fruit, slender
Flower: petals narrowly elliptic, white
Seeds 0.7–0.8 mm, compressed, with marginal ridge opposite an elongate depression; tubercles low, rounded
Chromosomes: 2n=20
Ecology: Uncommon. Disturbed areas
Elevation: < 850 m.
Bioregional distribution: Klamath Ranges, Cascade Range, Central Coast
Distribution outside California: native to c, e, s Europe

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bioregional map for HOLOSTEUM%20umbellatum%20subsp.%20umbellatum being generated

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